Stylommatophora

Figure L4.Labels. i Bulimus altoperuvianus Reeve, 1849 ii Bulimus amandus Pfeiffer, 1855 iii Bulimus ambustus Reeve, 1849 iv Bulimus andicola Pfeiffer, 1847 v Bulimus andoicus Morelet, 1863 vi Drymaeus angustus da Costa, 1906.

Figure 12.Genitalia. A, B Discartemon discadentus sp. n. paratype CUMZ 6209 A reproductive system, and B insertion of vas deferens into penial sheath C, D Discartemon hypocrites topotype CUMZ 6199 C reproductive system, and D insertion of vas deferens into penis sheath.

Figure 9.A–B Aspastus species A, iv Aspastus miltocheilus manugiensis (B. Rensch, 1934), paratype ZMB 78674 (H = 67.1) B, ii Aspastus miltocheilus paravicinii (B. Rensch, 1934), paratype ZMB 78795 (H = 55.9) C–E Eumecostylus species C, i Eumecostylus hargravesi (Cox, 1871), paratype of Placostylus (Placocharis) hargravesi aukiensis W.F. Clapp, 1923 ZMB 74853 (H = 51.1) D, v Eumecostylus uliginosus (Kobelt, 1891), syntype ZMB 48239 (H = 50.0) E, iii Eumecostylus phenax (W.F. Clapp, 1923), paratype ZMB 74851 (H = 81.3).

Figure 10.Bostryx species. A–B Bostryx voithianus (Pfeiffer, 1847), lectotype NHMUK 20100565.1 (H = 19.1) B sculpture on dorsal side of (pen)ultimate whorl C Bostryx simpliculus (Pfeiffer, 1855), lectotype NHMUK 1975340 (H = 18.9) D Bostryx turritus (Broderip in Broderip and Sowerby I 1832), possible syntype NHMUK 20100616 (H = 25.3) E Bostryx virgultorum (Morelet, 1863), paralectotype NHMUK 1893.2.4.179 (H = 27.8) F Bostryx woodwardi (Pfeiffer, 1857), lectotype NHMUK 1975334 (H = 31.0) G Bostryx andoicus (Morelet, 1863), lectotype NHMUK 1893.2.4.171 (H = 29.5) H Bostryx derelictus (Broderip in Broderip and Sowerby I 1832), probable syntype NHMUK 20100609 (H = 30.3). All enlarged.

Figure 1.Bostryx species. A Bostryx umbilicaris (Souleyet, 1842), paralectotype NHMUK 1854.7.24.359 (H = 13.3) B Bostryx moniezi (Dautzenberg, 1896), syntype NHMUK 1908.6.13.22 (H = 12.46) C–D Bostryx infundibulum (Pfeiffer, 1853), lectotype NHMUK 1975163 (H = 18.25) E Bostryx scalaricosta (Morelet, 1860), lectotype NHMUK 1893.2.4.1170 (H = 15.5) F Bostryx hamiltoni (Reeve, 1849), lectotype NHMUK 1849.5.14.53 (H = 17.0) G–H Bostryx lesueurianus (Morelet, 1860), syntype NHMUK 1893.2.4.1182 (H = 22.5) I Bostryx pygmaeus costatus Weyrauch, 1960, paratype NHMUK 1975357 (H = 8.45) J Bostryx imeldae costifer Weyrauch, 1960, paratype NHMUK 1975354 (H = 9.6) K Bostryx metagyra Pilsbry & Olsson, 1949, paratype NHMUK 20100630 (H = 7.1). All enlarged.

Figure 17.Internal sculpture of genitalia. A–E Discartemon nummus CUMZ 6208, from Patthalung A details of atrium surface B low magnification shows arrangement of penial hooks C high magnification of penial hooks D top view of penial hooks, white arrow indicate tip of hook, and E arrangement of vaginal folds F–I Discartemon discadentus sp. n., paratype CUMZ 6209 F details of atrium surface G high magnification of penial hooks with (inset) shows in low magnification H top view of penial hook, and I arrangement of reticulated vaginal folds.

Figure 16.A–B Bulimulus species A, i Bulimulus coriaceus (Pfeiffer, 1857) syntype ZMB 117767 (H = 12.9) B, ii Bulimulus monachus (Pfeiffer, 1857) syntype ZMB 117773 (H = 24.2) C, iii Drymaeus (Mesembrinus) jonasi (Pfeiffer, 1846) syntype ZMB 117769 (H = 21.5) D–F Cochlorina involuta (Martens, 1867), holotype ZMB 117768 (H = 15.2). All enlarged.

Figure L17.Labels. i Bulimus delumbis Reeve, 1849 ii Bulimus dentritis Morelet, 1863 iii Bulimus demotus Reeve, 1850 iv Bulimus denickei J.E. Gray, 1852 v Bulimus depictus Reeve, 1849 vi Bulimus depstus Reeve, 1849 vii Bulinus derelictus Broderip in Broderip and Sowerby I 1832.

Figure L33.Labels. i Bulimus (Otostomus) lamas Higgins, 1868 ii Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895 iii Bulimulus latecolumellaris Preston, 1909 iv Bulimus lattrei Pfeiffer in Philippi 1846 v Bulimulus laxostylus Rolle, 1904 vi Bulimus lesueureanus Morelet, 1860.

Figure 23.Approximate locations of the type locality of Discartemon species. (1) Discartemon discus, (2) Discartemon planus, (3) Discartemon sykesi, (4) Discartemon nummus, (5) Discartemon khaosokensis, (6) Discartemon discadentus sp. n., (7) Discartemon discamaximus sp. n., (8) Discartemon circulus sp. n., (9) Discartemon deprima sp. n., (10) Discartemon expandus sp. n., (11) Discartemon plussensis, (12) Discartemon hypocrites, (13) Discartemon leptoglyphus, (14) Discartemon platymorphus, (15) Discartemon afthonodontia sp. n., (16) Discartemon epipedis sp. n., (17) Discartemon flavacandida sp. n., (18) Discartemon lemyrei, (19) Discartemon roebeleni, (20) Discartemon collingei, (21) Discartemon stenostomus, (22) Discartemon sangkarensis, (23) Discartemon vandermeermohri, (24) Discartemon kotanensis sp. n., (25) Discartemon megalostraka sp. n., (26) Discartemon triancus sp. n., and (27) Discartemon conicus sp. n.

Figure 15.Bostryx (sensu lato) species A–C Bostryx stelzneri (Dohrn, 1875) A, i possible syntype of Bulimulus (Scutalus) stelzneri Dohrn, 1875 ZMB 34734 (H = 27.9) B, ii syntype of Bulimulus (Scutalus) conospirus Doering, 1879 ZMB 34721 (H = 21.9) C, iii syntype of Bulimulus (Scutalus) peristomatus Doering, 1879 ZMB 34723 (H = 26.5) D–E, iv Bostryx tortoranus (Doering, 1879) syntype ZMB 34718 (H = 21.9) F–G, v Bostryx tortoranus (Doering, 1879), syntype of Bulimulus (Bulimulus) monticola Doering, 1879 ZMB 34725 (H = 13.05).All enlarged.

Figure 24.Drymaeus species. A Drymaeus (Mesembrinus) flavidulus (E.A. Smith, 1877), lectotype NHMUK 1975134 (H = 21.4) B Drymaeus (Mesembrinus) fuscobasis (E.A. Smith, 1877), lectotype NHMUK 1975139 (H = 28.7) C Drymaeus (Mesembrinus) virginalis (Pfeiffer, 1856), probable syntype NHMUK 1975503 (H = 27.9) D Drymaeus (Mesembrinus) deshayesi (Pfeiffer, 1845), lectotype NHMUK 1975526 (H = 44.5) E Drymaeus (Mesembrinus) columbianus (Lea, 1838), lectotype of Bulimus gruneri Pfeiffer, 1846 NHMUK 20100563/1 (H = 27.8) F Drymaeus (Mesembrinus) rectilinearis (Pfeiffer, 1855), NHMUK 1975337 (H = 23.9) G Drymaeus (Mesembrinus) multilineatus (Say, 1825), paralectotype of Bulimus sisalensis Morelet, 1849 NHMUK 1893.2.4.1655 (H = 24.1) H Drymaeus (Mesembrinus) koppeli (Sowerby III, 1892), lectotype NHMUK 1907.11.21.133 (H = 25.5) I Drymaeus (Mesembrinus) hondurasanus (Pfeiffer, 1846), lectotype NHMUK 1975265 (H = 28.6) J Drymaeus (Mesembrinus) inglorius (Reeve, 1848), lectotype NHMUK 1975536 (H = 25.6) K Drymaeus (Mesembrinus) tripictus hoffmanni (Martens, 1893), holotype of Drymaeus prestoni cancellata da Costa, 1906 NHMUK 1907.11.21.13 (H = 18.1) L Drymaeus (Mesembrinus) gabbi (Angas, 1879), lectotype NHMUK 1879.7.22.23 (H = 20.6). All enlarged.

Figure 53.Drymaeus species. A–C Drymaeus (Drymaeus) lilacinus lilacinus (Reeve, 1849), probably syntype NHMUK 20100520 (H = 45.5) D–F Drymaeus (Drymaeus) lilacinus jansoni (Martens, 1893), lectotype NHMUK 1901.6.22.951 (H = 45.7) G–I Drymaeus (Drymaeus) malleatus (da Costa, 1898), holotype NHMUK 1907.11.21.130 (H = 34.5). All enlarged.

Figure 5.Shells of Group I: Discartemon discus-group. A, B Discartemon discamaximus sp. n. A holotype CUMZ 6245 with apertural dentition, and B paratype CUMZ 6005 C, D Discartemon circulus sp. n. C holotype CUMZ 6246 with apertural dentition, and D paratype CUMZ 3665 E, F Discartemon deprima sp. n. E holotype CUMZ 6247 with apertural dentition, and F paratype CUMZ 3573 G, H Discartemon expandus sp. n. G holotype CUMZ 6248 with apertural dentition, and H paratype CUMZ 3664.

Figure 17.Drymaeus (Mesembrinus) species A–B, i Drymaeus (Mesembrinus) bahamensis (Pfeiffer, 1862), syntype ZMB 25727 (H = 29.4) C–D, ii Drymaeus (Mesembrinus) flavidus (Menke, 1829) probable syntype ZMB 10338 (H = 23.4) E, iii Drymaeus (Mesembrinus) livescens (Pfeiffer, 1842), probable syntype ZMB 117771 (H = 22.5) F–G, vi Drymaeus (Mesembrinus) loxensis (Pfeiffer, 1846), syntype ZMB 117772 (H = 24.7) H, v Drymaeus (Mesembrinus) moussoni (Pfeiffer, 1853), syntype ZMB 117774 (H = 23.5) I, iv Drymaeus (Mesembrinus) translucens (Broderip in Broderip and Sowerby I, 1832), probable syntype ZMB 117778 (H = 13.3).

Figure L18.Labels. i Bulimus deshayesi Pfeiffer, 1845 ii Bulimus devians Dohrn, 1863 iii Bulinus discrepans Sowerby I, 1833 iv Bulimus dubius Pfeiffer, 1853 v Bulimus dombeyanus ‘Férussac’ Pfeiffer, 1846.

Figure L34.Labels. i Helix lichnorum d’Orbigny, 1835 ii Helix limonoica d’Orbigny, 1835 iii Bulimus lilacinus Reeve, 1849 iv Helix linostoma d’Orbigny, 1835 v Bulimus limensis Reeve, 1849 vi Bulimus liquabilis Reeve, 1848 vii Helix lithoica d’Orbigny, 1835 viii Bulimus lirinus Morelet, 1851 ix Bulimus lividus Reeve, 1850.

Figure 23.Approximate locations of the type locality of Discartemon species. (1) Discartemon discus, (2) Discartemon planus, (3) Discartemon sykesi, (4) Discartemon nummus, (5) Discartemon khaosokensis, (6) Discartemon discadentus sp. n., (7) Discartemon discamaximus sp. n., (8) Discartemon circulus sp. n., (9) Discartemon deprima sp. n., (10) Discartemon expandus sp. n., (11) Discartemon plussensis, (12) Discartemon hypocrites, (13) Discartemon leptoglyphus, (14) Discartemon platymorphus, (15) Discartemon afthonodontia sp. n., (16) Discartemon epipedis sp. n., (17) Discartemon flavacandida sp. n., (18) Discartemon lemyrei, (19) Discartemon roebeleni, (20) Discartemon collingei, (21) Discartemon stenostomus, (22) Discartemon sangkarensis, (23) Discartemon vandermeermohri, (24) Discartemon kotanensis sp. n., (25) Discartemon megalostraka sp. n., (26) Discartemon triancus sp. n., and (27) Discartemon conicus sp. n.

Figure 33.Spixia species A–B, i Spixia alvarezii (d’Orbigny, 1837), syntype of Bulimus (Odontostomus) leptodon Martens, 1875 ZMB 24077 (H = 22.2) C, ii Spixia martensii (Doering, 1875), paralectotype ZMB 28807 (H = 21.4) D, v Spixia multispirata (Doering, 1877), paralectotype ZMB 28509 (H = 17.4) E, iii Spixia philippii (Doering, 1875), syntype ZMB 28504 (H = 18.0) F, iv Spixia reticulata (Doering, 1877), paralectotype ZMB 28502 (H = 18.8) G, vi Spixia salinicola (Doering, 1877), paralectotype ZMB 28506 (H = 20.4) H, vii Spixia tumulorum (Doering, 1875), paralectotype ZMB 25739 (H = 13.6). All enlarged.

Figure 4.Bostryx species. A Bostryx rodriguezae Weyrauch, 1967, paratype NHMUK 1975355 (H = 25.5) B Bostryx rehderi Weyrauch, 1960, paratype NHMUK 1975356 (H = 10.45) C Bostryx devians (Dohrn, 1863), syntype NHMUK 1975339 (H = 14.9) D Bostryx mordani Breure, 1978, paratype NHMUK 1975266 (H = 22.5) E Bostryx agueroi Weyrauch, 1960, paratype NHMUK 1975333 (H = 21.1) F–H Bostryx orophilus (Morelet, 1860) F Bulimus albicolor Morelet, 1863, syntype NHMUK 1893.2.4.169 (H = 28.2) G Bulimus cercicola Morelet, 1863, syntype NHMUK 1893.2.4.175 (H = 18.7) H Bulimus orophilus Morelet, 1860, lectotype NHMUK 1893.2.4.188 (H = 20.8). All enlarged.

Figure 6.Bostryx species. A Bostryx acalles (Pfeiffer, 1853), possible syntype NHMUK 20100651 (H = 13.2) B Bostryx depstus (Reeve, 1849), lectotype NHMUK 1975318 (H = 18.5) C Bostryx conspersus (Sowerby I, 1833), probable syntype NHMUK 20100619 (H = 19.8) D Bostryx ?hennahi (J.E. Gray, 1830), lectotype of Helix cactorum d’Orbigny, 1835, NHMUK 1854.12.4.189 (H = 24.4) E–G Bostryx modestus (Broderip in Broderip and Sowerby I 1832) E Lectotype of Bulimus limensis Reeve, 1849, NHMUK 1975326 (H = 19.6) F syntype of Bulimus philippii Pfeiffer, 1842, NHMUK 1975348 (H = 24.3) G Bulinus modestus Broderip in Broderip and Sowerby I 1832, possible syntype NHMUK 20120232 (H = 31.6). All enlarged.

Figure 5.Shells of Group I: Discartemon discus-group. A, B Discartemon discamaximus sp. n. A holotype CUMZ 6245 with apertural dentition, and B paratype CUMZ 6005 C, D Discartemon circulus sp. n. C holotype CUMZ 6246 with apertural dentition, and D paratype CUMZ 3665 E, F Discartemon deprima sp. n. E holotype CUMZ 6247 with apertural dentition, and F paratype CUMZ 3573 G, H Discartemon expandus sp. n. G holotype CUMZ 6248 with apertural dentition, and H paratype CUMZ 3664.

Figure 19.A–C Liguus fasciatus (Müller, 1774) A, v Probable syntype of Achatina blainiana Poey, 1853 ZMB 117781 (H = 39.0) B, i Paratype of Liguus fasciatus archeri Clench, 1934 ZMB 78796 (H = 51.9) C, ii Paratype of Liguus crenatus barbouri Clench, 1929 ZMB 74876 (H = 46.3) D–E, iii Orthalicus elegans Rolle, 1895, lectotype ZMB 47655 (H = 61.2) F–G, iv Orthalicus nobilis Rolle, 1895, lectotype ZMB 47656 (H = 58.8).

Figure L18.Labels. i Bulimus deshayesi Pfeiffer, 1845 ii Bulimus devians Dohrn, 1863 iii Bulinus discrepans Sowerby I, 1833 iv Bulimus dubius Pfeiffer, 1853 v Bulimus dombeyanus ‘Férussac’ Pfeiffer, 1846.