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↑Forrest, Laura L.; Christine E. Davis; David G. Long; Barbara J. Crandall-Stotler; Alexandra Clark; Michelle L. Hollingsworth (2006). “Unraveling the evolutionary history of the liverworts (Marchantiophyta): multiple taxa, genomes and analyses”. 《The Bryologist》 109 (3): 303–334. doi:10.1639/0007-2745(2006)109[303:UTEHOT]2.0.CO;2.