dcsimg

Description

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Trees , to 30 m; crowns oblong. Bark gray, deeply fissured with broad, flattened ridges. Wood hard. Branches short-spreading, young branches pubescent, old-growth with 3-5 prominent, irregular, corky wings; twigs reddish, pubescent. Buds brown, ovoid, acute, pubescent; scales brown, pilose on outer surface, ciliate on margins. Leaves: petiole ca. 5 mm, pubescent. Leaf blade obovate to oblong-oval, (2.5-)9-11(-16) × 2.5-5 cm, base oblique, margins doubly serrate, apex short-acuminate; surfaces abaxially white-pubescent, pubescence not tufted in axils of veins, adaxially dark green, usually glabrous, sometimes scabrous. Inflorescences racemose cymes, long-pendulous, (7-)10(-13)-flowered, to 5 cm; pedicel 0.5-1 cm. Flowers: calyx deeply lobed, divided nearly to middle, lobes 7-8; stamens 5-8; anthers dark purple; stigmas greenish, pubescent. Samaras elliptic to oval, 1.5-2.2 cm, narrowly winged, pubescent, margins short-ciliate, apex shallowly notched. Seeds inflated, not thickened. 2 n = 28.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Distribution

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Ont.; Ark., Ill., Ind., Iowa, Kans., Ky., Mich., Minn., Mo., Nebr., N.H., N.J., N.Y., Ohio, S.Dak., Tenn., Vt., W.Va., Wis.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flowering/Fruiting

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Flowering spring.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Habitat

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Rocky slopes, limestone outcrops, rich woods, flood plains, stream banks; 30-900m.
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Synonym

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Ulmus racemosa Thomas 1831, not Borkhausen 1800
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Associated Forest Cover

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Rock elm is a minor component in two forest cover types: Sugar Maple-Beech-Yellow Birch (Society of American Foresters Type 25) and Black Ash-American Elm-Red Maple (Type 39). In addition to type species, other important associates include white ash (Fraxinus americana), black cherry (Prunus serotina), basswood (Tilia spp.), northern red oak (Quercus rubra), American elm (Ulmus americana), eastern hophornbeam (Ostrya virginiana), and eastern hemlock (Tsuga canadensis).

Some of the woody shrubs commonly associated with rock elm include prickly ash (Zanthoxylum americanum), beaked hazel (Corylus cornuta), blackberry and raspberry (Rubus spp.), dogwoods (Cornus spp.), gooseberry (Ribes spp.), Atlantic leatherwood (Dirca palustris), bittersweet (Celastrus scandens), Virginia creeper (Parthenocissus quinquefolia), grape (Vitis spp.), hawthorn (Crataegus spp.), American and redberry elder (Sambucus canadensis and S. pubens), and nannyberry (Viburnum lentago).

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Climate

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The climatic conditions associated with the distribution of rock elm can be characterized as continental, with cold winters and warm summers. Within the species range, a maximum summer temperature of 38° C (100° F) and a minimum winter temperature of -34° C (-30° F) are common.

Annual precipitation is 640 mm (25 in) in the western part of the range and 1270 mm (50 in) in the extreme southern and eastern parts. At least half of this precipitation occurs during the growing season. Snowfall averages from 50 to 200 cm (20 to 80 in), depending on geographic location.

The frost-free period averages 100 days in the north and 200 days in the south. Rock elm grows best where the frost-free period is from 120 to 160 days.

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Damaging Agents

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Nearly all native North American elm species are susceptible to Dutch elm disease (Ceratocystis ulmi) (6,13) and isolates of C. ulmi have been obtained from rock elm logs (5). It is likely that Dutch elm disease will greatly reduce the number of rock elm..

A seed-borne fungus (Gleosporium ulmicolum) has been reported for rock elm but few of the fungi that are able to invade the fruits and seeds of North American hardwoods are thought to be pathogens that reduce germination or weaken seedlings (1).

Although rock elm has not been listed as a particular host for specific insects, undoubtedly it is host to the various borers, defoliators, and sucking insects that attack American elm.

Throughout the range of rock elm, killing frosts are common during the flowering period and subfreezing temperatures may prevent seed development in some years.

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Flowering and Fruiting

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Rock elm flowers appear 2 weeks before the leaves at any time from March to May, depending on locality and site. The perfect flowers are protandrous, that is, the male elements of the flower develop 2 to 4 days before the female elements are receptive (6). Female flowers are receptive for only a few days.

The hairy fruit has a broad wing from 13 to 25 mm (0.5 to 1 in) long and matures during May or June. Clean, fully ripened, unwinged seeds number from 11,000 to 19,800/kg (5,000 to 9,000/lb). Seeds germinate soon after they ripen.

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Genetics

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In a study of compatibility and crossability in Ulmus (11), the form of dichogamy (protandry or protogyny) correlated with the compatibility between the different species. Ulmus thomasii is a protandry species and is compatible with two other protandry species-U. pumila and U. laevis. Ulmus thomasii is also self-fertile.

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Growth and Yield

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Few species have rock elm's capacity for recovering from prolonged suppression. An analysis of 153 trees indicated that a large percentage had survived under suppression for 50 years or more. This capacity makes it difficult to correlate diameter and age (9):



Crown class D.b.h. class Dominant Codominant Intermediate Suppressed 2.5 cm or 1 in 14 22 30 48 7.6 cm or 3 in 26 50 64 99 12.7 cm or 5 in 39 72 97 -- 17.8 cm or 7 in 51 93 -- -- 22.9 cm or 9 in 63 -- -- --

The average number of rings per 2.5 cm (1 in) of radius by crown class was about 50 for suppressed, 30 to 40 for intermediate, 20 to 30 for codominants, and 10 to 20 for dominants.

On average to better sites, mature rock elm may reach 27 in (90 ft) in total height and 61 cm (24 in) in d.b.h. (12). In virgin hardwood stands in the east and north, 27 to 30 in (90 to 100 ft) heights and 91 to 152 cm (36 to 60 in) in d.b.h. have been recorded (10). Much smaller trees occur along river bluffs, on limestone outcrops, or other sites with thin soil mantles. Rock elm may live 250 to 300 years.

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Reaction to Competition

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Rock elm is considered shade tolerant in the seedling-sapling stage and often recovers successfully after long periods of suppression at these stages. As the tree grows older, however, it apparently becomes more light demanding. Overall, the species is classed as intermediate in tolerance to shade (10).

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Rooting Habit

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No information available.

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Seed Production and Dissemination

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Trees 20 years old produce viable seeds, but maximum yields are from trees 45 to 125 years old. Good crops occur every 3 or 4 years. Ripe seeds are dispersed as the leaves become fully expanded, which is usually 2 or 3 weeks later than the time of seed drop for American elm (9).

Although the thin, hair-fringed, winged samaras seem adapted to wind dispersal, seeds are generally carried no more than 40 to 45 m (100 to 150 ft) from the parent tree. The fact that rock elm grows as scattered individuals, often several miles from the nearest seed source, suggests that birds and small mammals play a role in dissemination. The large but very light seeds are also buoyant and water can carry them long distances. As a result, seeds often are concentrated along the banks of streams and lakes.

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Seedling Development

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Rock elm seeds germinate within a week or two after dispersal if moisture conditions are favorable. In a germination test, 90 to 100 percent of mature seeds were viable (2). Viability was not significantly different between seeds from different trees, between seeds with wings or without wings, or between seeds with seed coats or without seed coats. When germinated in a petri dish, radicles of the viable seeds emerged within 2 or 3 days and were 2.5 to 3.8 cm (1 to 1.5 in) long by the end of the fifth day Germination is epigeal. The cotyledons began to open about the fifth or sixth day. Under favorable conditions, rock elm seedlings are from 5 to 8 cm (2 to 3 in) tall by the end of the first summer.

Despite its high seed viability, rock elm regenerates poorly (2,9). Germination tests failed when mineral soil and equal volumes of peat moss, sand, and mineral soil were used for planting media, but 70 to 80 percent emergence was obtained in flats using peat moss. Another factor affecting seedling establishment is the persistence of dormant terminal buds. Emergent seedlings rarely develop more than a single pair of true leaves during the first growing season due to this dormancy. Observations on more than 200 seedlings indicated that only 1 percent broke dormancy long enough to develop an additional internode and a second pair of true leaves.

This species appears to be shade tolerant during the seedling stage (10). However, under field conditions with competition, 1.5-0 nursery stock averaged only 27 cm (10.6 in) in height 5 years after planting and only 52 cm (20.4 in) 10 years after planting in northern Wisconsin. In the same study, survival ranged from 85 percent at the end of the 1st year to 32 percent at the end of the 10th year.

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Soils and Topography

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Rock elm is most frequent in lower Michigan, Wisconsin, and southern Ontario, and it is regularly found on moist but well-drained sandy loam, loam, or silt loam soils in mixture with other hardwoods. In Wisconsin, rock elm is most frequent in the southern wet-mesic forest (7). Although rock elm often grows on rocky ridges, limestone outcroppings, and streambanks, the highest quality sawtimber is found on deeper loamy soils.

The major soil orders associated with the distribution of rock elm are the Mollisols, Alfisols, and the Spodosols. Most common are the Hapludalfs (Gray-Brown Podzolic soils) within the Udalfs suborder of the Alfisols. Soil pH ranges from slightly alkaline or neutral to strongly acid.

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Special Uses

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The seeds and buds of rock elm are eaten by deer, rabbits, squirrels, and a variety of birds. Small mammals such as chipmunks, ground squirrels, and mice apparently relish the filbertlike flavor of rock elm seed and frequently eat the major part of the crop.

Rock elm wood has long been valued for its exceptional strength and superior quality (3,8). For this reason rock elm has been drastically overcut in many localities. The wood is stronger, harder, and stiffer than any of the other commercial species of elms. It is highly shock resistant and has excellent bending qualities which make it good for bent parts of furniture, crates and containers, and a base for veneer. Much of the old-growth was exported for ship timbers. Currently, the highest quality sawtimber is found in north-central Wisconsin, lower Michigan, and southeastern Ontario.

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Vegetative Reproduction

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Rock elm regenerates vegetatively from root suckers and stump sprouts (10), but vegetative reproduction in the field is uncommon.

Most elms are considered difficult to root by means of cuttings. However, leaf-bud cuttings, consisting of leaf blade, axillary bud, and a shield of stem tissue, treated with a growth hormone and held under constant mist on a rooting medium of sand or mica, produced satisfactory results for several species of elm including rock elm (5).

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Brief Summary

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Ulmaceae -- Elm family

T. R. Crow

Rock elm (Ulmus thomasii), often called cork elm because of the irregular thick corky wings on older branches, is a medium-sized to large tree that grows best on moist loamy soils in southern Ontario, lower Michigan, and Wisconsin. It may also be found on dry uplands, especially rocky ridges and limestone bluffs. On good sites, rock elm may reach 30 m (100 ft) in height and 300 years of age. It is always associated with other hardwoods and is a valued lumber tree. The extremely hard, tough wood is used in general construction and as a veneer base. Many kinds of wildlife consume the abundant seed crops.

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Distribution

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Rock elm is most common to the Upper Mississippi Valley and lower Great Lakes region. The native range includes portions of New Hampshire, Vermont, New York, and extreme southern Quebec; west to Ontario, Michigan, northern Minnesota; south to southeastern South Dakota, northeastern Kansas, and northern Arkansas; and east to Tennessee, southwestern Virginia, and southwestern Pennsylvania. Rock elm also grows in northern New Jersey.


-The native range of rock elm.


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Ulmus thomasii

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Ulmus thomasii, the rock elm[3] or cork elm (or orme liège in Québec), is a deciduous tree native primarily to the Midwestern United States. The tree ranges from southern Ontario and Quebec, south to Tennessee, west to northeastern Kansas, and north to Minnesota.[4]

Etymology

The tree was named in 1902 for David Thomas, an American civil engineer who had first named and described the tree in 1831 as Ulmus racemosa.[5]

Description

Ulmus thomasii grows as a tree from 15–30 m (50–100 ft) tall, and may live for up to 300 years. Where forest-grown, the crown is cylindrical and upright with short branches, and is narrower than most other elms.[6] Rock elm is also unusual among North American elms in that it is often monopodial.[7] The bark is grey-brown and deeply furrowed into scaly, flattened ridges. Many older branches have 3–4 irregular thick corky wings. It is for this reason the rock elm is sometimes called the cork elm.[8]

The leaves are 5–10 cm (2–4 in) long and 2–5 cm (34–2 in) wide, oval to obovate with a round, symmetrical base and acuminate apex. The leaf surface is shiny dark green, turning bright yellow in autumn; the underside is pubescent. The perfect apetalous, wind-pollinated flowers are red-green and appear in racemes up to 40 mm (2 in) long two weeks before the leaves from March to May, depending on the tree's location. The fruit is a broad ovate samara 13–25 mm (123132 in) long covered with fine hair, notched at the tip, and maturing during May or June to form drooping clusters at the leaf bases.[9]

Although U. thomasii is protandrous, levels of self-pollination remain high.[10]

Ecology

Ulmus thomasii is moderately shade-tolerant.[11] Its preferred habitat is moist but well-drained sandy loam, loam, or silt loam soil, mixed with other hardwoods. However, it also grows on dry uplands, especially on rocky ridges and limestone bluffs.

Pests and diseases

Like most North American elms, U. thomasii is very susceptible to Dutch elm disease.

Cultivation

There are no known cultivars of Ulmus thomasii, nor is it known to be any longer in commerce. It appeared in some US nursery catalogues in the early 20th century.[12][13][14] The species is occasionally grown beyond its native range as a specimen tree in botanical gardens and arboreta, for example in northwestern Europe, but not commonly cultivated in northern Europe, being unsuited to the region's more temperate, maritime climate. However, the tree was propagated and marketed in the UK by the Hillier & Sons nursery, Winchester, Hampshire, from 1965 to 1977, during which time 49 were sold.[15][16]

Ulmus thomasii was crossed experimentally with Japanese elm (U. davidiana var. japonica) at the Arnold Arboretum in Massachusetts, but no clones were released to commerce.[10] Seedlings arising from crossings with Siberian elm (U. pumila) at the Lake States Forestry Experimental Station in the 1950s all perished,[17] a classic case of hybrid lethality.[18]

Notable trees

The US National Champion, measuring 100 ft (30 m) high in 1989, grows in Cass County, Michigan.[19]

Uses

The wood of the rock elm is the hardest and heaviest of all elms, and where forest-grown remains comparatively free of knots and other defects. It is also very strong and takes a high polish, and consequently was once in great demand in America and Europe for a wide range of uses, notably boatbuilding, furniture, agricultural tools, and musical instruments.

Much of the timber's strength is derived from the tight grain arising from the tree's very slow rate of growth, the trunk typically increasing in diameter by less than 2 mm (332 in) a year. Over 250 annual growth rings were once counted in a log 24 cm (9+12 in) square being sawn for gunwales in an English boatyard, while a tree once grown at Kew Gardens, London, attained a height of only 12 m (39 ft) in 50 years.[20]

Accessions

North America
Europe

References

Wikimedia Commons has media related to Ulmus thomasii.
  1. ^ Stritch, L. (2018). "Ulmus thomasii". IUCN Red List of Threatened Species. 2018: e.T61967392A61967401. doi:10.2305/IUCN.UK.2018-1.RLTS.T61967392A61967401.en. Retrieved 18 November 2021.
  2. ^ "Ulmus thomasii". Tropicos. Missouri Botanical Garden.
  3. ^ BSBI List 2007 (xls). Botanical Society of Britain and Ireland. Archived from the original (xls) on 2015-06-26. Retrieved 2014-10-17.
  4. ^ "Ulmus Thomasii Range Map" (PDF). United States Geological Survey. Retrieved 2008-03-02.
  5. ^ This name had been used in 1800 for a different species of elm, hence the need for the later renaming that honored Thomas.
  6. ^ Photographs of mature Rock Elm showing narrow profile: Virginia Tech Department of Forest Resources [1], Natural Resources of Canada, tidcf.nrcan.gc.ca [2] Archived 2016-08-02 at the Wayback Machine [3]
  7. ^ Bean, W. J. (1981). Trees and shrubs hardy in Great Britain, 7th edition. Murray, London.
  8. ^ Photograph of corky ridges of Rock Elm branches, Michigan State University Plant Encyclopedia [4]
  9. ^ White, J & More, D. (2003). Trees of Britain & Northern Europe. Cassell's, London.
  10. ^ a b Hans, A. S. (1981). "Compatibility and Crossability Studies in Ulmus". Silvae Genetica. 30: 4–5.
  11. ^ "Forestry".
  12. ^ Kelsey, Frederick W., Choice Trees, cat. 55, N.Y. 1906, p.20
  13. ^ Griffing's tree & plant book, 1929; Griffing's Interstate Nurseries, C.M. Griffing & Company;p.29
  14. ^ Griffing's tree & plant book, 1930; Griffing's Interstate Nurseries; p 20
  15. ^ Hillier & Sons (1977). Catalogue of Trees & Shrubs. Hillier, Ampfield, UK.
  16. ^ Hillier & Sons Sales inventory 1962 to 1977 (unpublished).
  17. ^ Sholtz, H. F. (1957). Rock Elm (Ulmus thomasii). Lake States Forest Experimental Station Paper 47:16.
  18. ^ Mino, Masanobu; Maekawa, Kenji; Ogawa, Ken'Ichi; Yamagishi, Hiroshi; Inoue, Masayoshi (2002). "Cell Death Processes during Expression of Hybrid Lethality in Interspecific F1 Hybrid between Nicotiana gossei Domin and Nicotiana tabacum". Plant Physiology. 130 (4): 1776–1787. doi:10.1104/pp.006023. PMC 166689. PMID 12481061.
  19. ^ "Rock Elm (Ulmus thomasii)". The 2012 National Register of Big Trees. American Forests. 2012.
  20. ^ Elwes, H. J. & Henry, A. (1913). The Trees of Great Britain & Ireland. Vol. VII. 1848–1929. Republished 2004 Cambridge University Press, ISBN 978-1-108-06938-0
  21. ^ "The National Botanic Gardens of Ireland".

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Ulmus thomasii: Brief Summary

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Ulmus thomasii, the rock elm or cork elm (or orme liège in Québec), is a deciduous tree native primarily to the Midwestern United States. The tree ranges from southern Ontario and Quebec, south to Tennessee, west to northeastern Kansas, and north to Minnesota.

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