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Associations

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Foodplant / gall
Aceria erinea causes gall of live leaf of Juglans regia

In Great Britain and/or Ireland:
Foodplant / pathogen
Armillaria mellea s.l. infects and damages Juglans regia

Foodplant / shot hole causer
epiphyllous, almost entirely immersed pycnidium of Ascochyta coelomycetous anamorph of Ascochyta juglandis causes shot holes on leaf of Juglans regia

Foodplant / saprobe
fruitbody of Auricularia auricula-judae is saprobic on wood of Juglans regia

Foodplant / sap sucker
Callaphis juglandis sucks sap of Juglans regia

Foodplant / saprobe
colony of Chalaropsis dematiaceous anamorph of Ceratocystis paradoxa is saprobic on husk of Juglans regia

Foodplant / sap sucker
Chromaphis juglandicola sucks sap of Juglans regia

Foodplant / saprobe
gregarious, covered, plurilocular, stroma of Cytospora coelomycetous anamorph of Cytospora juglandina is saprobic on dead twig of Juglans regia
Remarks: season: 5

Foodplant / saprobe
bracket of Daedaleopsis confragosa is saprobic on dead wood of Juglans regia
Other: minor host/prey

Foodplant / saprobe
covered pycnidium of Phomopsis coelomycetous anamorph of Diaporthe juglandina is saprobic on dead bark of Juglans regia
Remarks: season: 8

Foodplant / saprobe
immersed, erumpent, conical, surrounded by laciniae, verruciform, paucilocellate, 1-2mm broad stroma of Fusicoccum coelomycetous anamorph of Fusicoccum juglandinum is saprobic on twig of Juglans regia

Foodplant / parasite
fruitbody of Ganoderma applanatum parasitises live trunk of Juglans regia
Other: minor host/prey

Foodplant / parasite
fruitbody of Inonotus hispidus parasitises live trunk of Juglans regia
Other: major host/prey

Foodplant / saprobe
superficial pycnidium of Aposphaeria coelomycetous anamorph of Melanomma pulvis-pyrius is saprobic on dry, hard, decorticate branch wood of Juglans regia
Remarks: season: 9-5

Foodplant / spot causer
hypophyllous, pulvinate, erumpent acervulus of Microstroma coelomycetous anamorph of Microstroma juglandis causes spots on live leaf of Juglans regia
Remarks: season: 7

Foodplant / pathogen
Tubercularia anamorph of Nectria cinnabarina infects and damages branch of Juglans regia
Remarks: season: 1-12

Foodplant / saprobe
fruitbody of Perenniporia fraxinea is saprobic on live trunk (base) of Juglans regia
Other: minor host/prey

Foodplant / saprobe
fruitbody of Phanerochaete sordida is saprobic on dead, decayed wood of Juglans regia
Other: minor host/prey

Foodplant / saprobe
stalked, clustered basidiocarp of Phleogena faginea is saprobic on dead, fallen trunk of Juglans regia
Remarks: season: 10-2

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Comments

provided by eFloras
The English or Persian walnut is found wild and cultivated in the Himalayas from 1000—3300 m alt. s. m. Ssp. fallax (Dode) Popov in Bull. App. Bot. 22, 3:204. 1929 (Juglans fallax Dode in Bull. Soc. Dendr. Fr. 89. 1906) is probably the form that occurs here. Valued for its wood and edible fruit; the wood is excellent for furniture, carving and for gun stocks; the bark is good for the gums and in the local market it is sold under the name of ‘dandasa’. It is also used as a vermi¬fuge and for staining; the leaves are used as fodder. The seed yields an oil used in cooking. The ‘kaghzi’ variety of walnut (Juglans doucloxiana of Dode) is valued for its thin shelled edible fruit.
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bibliographic citation
Flora of Pakistan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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S. I. Ali & M. Qaiser
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Comments

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Commonly cultivated in China from 23-42° N for its edible, oily nuts and hard, fine grained wood. Juglans regia has a very long history of cultivation in China and elsewhere; as a result, there are many cultivars, including five Chinese taxa that L.-A. Dode (Bull. Soc. Dendrol. France 2: 67-98. 1906), recognized on the basis of differences in shell thickness, size, etc.
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bibliographic citation
Flora of China Vol. 4: 282 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Description

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Deciduous tree up to 25 m tall. Young shoots tomentose. Leaves impa¬ripinnate, 17-40 cm long; leaflets 5-9, softly tomentose, opposite to sub-opposite, 7-20 cm long, 3-8 cm broad, ovate to elliptic-ovate, acute to acuminate, glab¬rescent to pubescent on nerves beneath; petiolule 2-4 mm long. Male catkins 6-12 cm long, lateral; bract c. 2 mm long; bracteoles 2, c. 3 nun long, ovate to obovate, pubescent; tepals 4, ovate, c. 2 mm long; stamens 10-20, subsessile; anthers 2 mm long, irregularly apiculate, basifixed, dehiscing longitudinally. Female flowers 1-3, terminal on short spikes; involucral tube of fused bract and bracteoles, c. 3.5 mm long, tomentose, glandular, obscurely 4-toothed and irregular at the margin; tepals 4, linear, 2.5-4 mm long, alternating with the teeth; margin sparsely pubescent; ovary c. 3.5 mm long, ovoid, inferior; style c. 2 mm long; stigmas 2, recurved, plumose to fimbriate, exserted. Drupe up to 5 cm long, ovoid to subglobose; epicarp green, glandular; endocarp 2-valved; seed 2 to 4-lobed at the base.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Description

provided by eFloras
Trees to 25 m tall. Leaves 25-30 cm; petiole 5-7 cm; petiole and rachis glabrescent, without glandular hairs; leaflets (3 or)5-9, entire on mature trees, sometimes obscurely serrulate on young plants; lateral leaflets subsessile or petiolule 1-2 mm, blade elliptic-ovate to long elliptic, 6-15 × 3-6 cm, abaxially glabrous except for tufts of hairs in vein axils, without glandular hairs, base oblique, subrounded, apex obtuse or acute to shortly acuminate; terminal petiolule 2.5-6 cm. Male spike 5-10(-15) cm. Stamens 6-30(-40). Fruiting spike usually with 1-3(-38) nuts. Nuts subglobose, 4-6 cm; husk glabrous, irregularly dehiscent; shell thick except in commercial varieties, wrinkled. Fl. Apr-May, fr. Oct. 2n = 32.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of China Vol. 4: 282 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Distribution

provided by eFloras
Distribution: C. America, SE. Europe, Caucasus, Syria, N. Iran, Afghanistan, W. Pakistan, Tibet, Nepal, W. China and Upper Burma.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Distribution

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Widely distributed in C, E, NW, and SW China [SW Asia to Himalayas, SE Europe]
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 282 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Flower/Fruit

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Fl. Per. February-April.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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eFloras.org
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Habitat

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Mountain slopes; 500-1800(-4000) m.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 282 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Synonym

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Juglans duclouxiana Dode; J. fallax Dode; J. kamaonia (C. de Candolle) Dode; J. orientis Dode; J. regia var. sinensis C. de Candolle; J. sinensis (C. de Candolle) Dode.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 282 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Derivation of specific name

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regia: splendid, royal
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Mark Hyde, Bart Wursten and Petra Ballings
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Hyde, M.A., Wursten, B.T. and Ballings, P. (2002-2014). Juglans regia L. Flora of Zimbabwe website. Accessed 28 August 2014 at http://www.zimbabweflora.co.zw/cult/species.php?species_id=166440
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Mark Hyde
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Bart Wursten
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Petra Ballings
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Juglans regia

provided by wikipedia EN

Juglans regia, the Persian walnut, English walnut, Carpathian walnut, Madeira walnut,[2] or especially in Great Britain, common walnut,[1] is an Old World walnut tree species native to the region stretching from the Balkans eastward to the Himalayas and southwest China. It is widely cultivated across Europe and the Caucasus.

It is the origin of cultivated varieties which produce the edible walnut, consumed around the world. China is the major commercial producer of walnuts.

Description

Juglans regia is a large deciduous tree, attaining heights of 25–35 metres (80–120 feet), and a trunk up to 2 m (6 ft) in diameter, commonly with a short trunk and broad crown.

The bark is smooth, olive-brown when young and silvery-grey on older branches, and features scattered broad fissures with a rougher texture. Like all walnuts, the pith of the twigs contains air spaces; this chambered pith is brownish in color. The leaves are alternately arranged, 25–40 cm (10 to 16 in) long, odd-pinnate with 5–9 leaflets, paired alternately with one terminal leaflet. The largest leaflets are the three at the apex, 10–18 cm (4 to 7 in) long and 6–8 cm (2 to 3 in) broad; the basal pair of leaflets are much smaller, 5–8 cm (2 to 3 in) long, with the margins of the leaflets entire. The male flowers are in drooping catkins 5–10 cm (2 to 4 in) long, and the female flowers are terminal, in clusters of two to five, ripening in the autumn into a fruit with a green, semifleshy husk and a brown, corrugated nut. The whole fruit, including the husk, falls in autumn; the seed is large, with a relatively thin shell, and edible, with a rich flavour.

Etymology

Taxonomic keys

The Latin name for the walnut was nux Gallica, "Gallic nut";[3] the Gaulish region of Galatia in Anatolia lies in highlands at the western end of the tree's presumed natural distribution.

For the etymology and meaning of the word in English and other Germanic languages, see "walnut".

"Walnut" does not distinguish the tree from other species of Juglans. Other names include common walnut in Britain; Persian walnut in South Africa[4] and Australia;[5] and English walnut in North America and Great Britain,[6] New Zealand,[7] and Australia,[5] the latter name possibly because English sailors were prominent in Juglans regia nut distribution at one time.[8] Alternatively, Walter Fox Allen stated in his 1912 treatise What You Need to Know About Planting, Cultivating and Harvesting this Most Delicious of Nuts:[9] "In America, it has commonly been known as English walnut to distinguish it from our native species."

In the Chinese language, the edible, cultivated walnut is called 胡桃 (hú táo in Mandarin), which means literally "Hu peach", suggesting the ancient Chinese associated the introduction of the tree into East Asia with the Hu barbarians of the regions north and northwest of China. In Mexico, it is called nogal de Castilla,[10] suggesting the Mexicans associated the introduction of the tree into Mexico with Spaniards from Castile (as opposed to the black walnuts native to North America).

The Old English term wealhhnutu is a late book-name (Old English Vocabularies, Wright & Wulker), so the remark that the Anglo-Saxons inherited the walnut tree from the Romans does not follow from this name. Old English: walhhnutu is wealh (foreign) + hnutu (nut). Etymologically it "meant the nut of the Roman lands (Gaul and Italy) as distinguished from the native hazel" according to the Oxford English Dictionary.

Distribution and habitat

Walnut tree - Juglans regia L. Claimed to be the oldest walnut tree in the world. Near Khotan, Xinjiang, China, in 2011
In August, Czech Republic

Original habitat

One of the centers of origin and diversity of Juglans regia is Iran.[11] However, as with other old and widespread cultivated plants, it is not easy to reconstruct the original distribution and determine the borders of the past natural ranges. There are many reports concerning the earliest fossil pollen and nuts of J. regia, and the conclusions that various authors draw are somewhat contradictory. Taken together these finds suggest that J. regia possibly survived the last glaciations in several refugia, as the compilation of the data shows most likely southern Europe, the Near East, China, and the Himalaya.[12]

The largest forests are in Kyrgyzstan, where trees occur in extensive forests at 1,000–2,000 metres (3,300–6,600 ft) altitude,[13] notably at Arslanbob in Jalal-Abad Province.

Introduction around the world

In the fourth century BC, Alexander the Great introduced this "Persian nut" (Theophrastus' καρυα ή Περσική[14]) in Macedonian, Ancient Greek ancestral forms with lateral fruiting from Iran and Central Asia. They hybridized with terminal-bearing forms to give lateral-bearing trees with larger fruit. These lateral-bearers were spread in southern Europe and northern Africa by Romans. Recent prospections in walnut populations of the Mediterrean Basin allowed to select interesting trees of this type. In the Middle Ages, the lateral-bearing character was introduced again in southern Turkey by merchants travelling along the Silk Road. J. regia germplasm in China is thought to have been introduced from Central Asia about 2,000 years ago, and in some areas has become naturalized. Cultivated J. regia was introduced into western and northern Europe very early, in Roman times or earlier, and to the Americas in the 17th century, by English colonists. Important nut-growing regions include California in the United States; France, Serbia, Greece, Bulgaria, Romania and Hungary in Europe; China in Asia; Baja California and Coahuila in Mexico; and Chile in Latin America. Lately, cultivation has spread to other regions, such as New Zealand and the southeast of Australia.[15] It is cultivated extensively from 30° to 50° of latitude in the Northern Hemisphere and from 30° to 40° in the Southern Hemisphere. Its high-quality fruits are eaten both fresh or pressed for their richly flavored oil; numerous cultivars have been selected for larger nuts with thinner shells.

Ecology

It tends to grow taller and narrower in dense forest competition. It is a light-demanding species, requiring full sun to grow well.

Juglans regia is infested by Rhagoletis juglandis, commonly known as the walnut husk fly, which lays its eggs in the husks of walnut fruit.

Other plants often will not grow under walnut trees because the fallen leaves and husks contain juglone, a chemical which acts as a natural herbicide. Horses that eat walnut leaves may develop laminitis, a hoof ailment.

Cultivation

Walnuts

Walnut trees grow best in rich, deep soil with full sun and long summers, such as the California central valley. Juglans hindsii and J. hindsii × J. regia are often used as grafting stock for J. regia.[17] Mature trees may reach 15 m (50 ft) in height and width, and live more than 200 years, developing massive trunks more than 2.4 m (8 ft) thick.

Cultivars

Walnut cultivars include:

  • Purpurea
  • Broadview
  • Cascade
  • Allegheny
  • Bedco 1
  • Coble #2
  • Hansen
  • Kaiser
  • KY Giant
  • Lake
  • McKinster
  • Somers
  • Utah Giant
  • Colby
  • Greenhaven
  • Reda
  • Shiawassee
  • Perry
  • S-1
  • Looking Glass
  • China-B
  • Champion
  • Northern Prize
  • Placentia

Pests

Particular cultivars of J. regia may be more infested by R. juglandis than others because of varying walnut husk softness or thickness. 'Eureka', 'Klondike', 'Payne', 'Franquette' and 'Ehrhardt' cultivars are among the most susceptible to infestation.[18]

Production

In 2019, world production of shelled walnuts was 4.5 million tonnes, led by China with 56% of the total harvested (table). Secondary producers were the United States and Iran.

Toxicity

Allergy

Walnuts and other tree nuts are food allergen sources having potential to cause life-threatening, IgE-mediated allergic reactions in some individuals.[19][20]

Uses

Nutrition

English walnut kernels are 4% water, 65% fat, 15% protein, and 14% carbohydrates (table). In a 100 gram reference amount providing 654 calories, the kernels supply several dietary minerals in rich content (20% or more of the Daily Value, DV), including manganese (162% DV), phosphorus (49% DV), magnesium (45% DV), zinc (33% DV), and iron (22% DV) among others, the B vitamins, B6 (42% DV), thiamine (30% DV), and folate (25% DV), and dietary fiber (table).

One study of various cultivars of J. regia in Turkey showed the fatty acid composition included about 6% palmitic acid, 3% stearic acid, 30% oleic acid, 50% linoleic acid, and 9% linolenic acid.[21]

Wood

Walnut heartwood is a heavy, hard, open-grained hardwood. Freshly cut live wood may be Dijon-mustard colour, darkening to brown over a few days. The dried lumber is a rich chocolate-brown to black, with cream to tan sapwood, and may feature unusual figures, such as "curly", "bee's wing", "bird's eye", and "rat tail", among others. It is prized by fine woodworkers for its durability, lustre and chatoyance, and is used for high-end flooring, guitars, furniture, veneers, knobs and handles as well as gunstocks.

Other uses

The Native American Navajo tribe has been documented using the hulls of the nut to create a brown dye.[22]

Culture

In Skopelos, a Greek island in the Aegean Sea, local legend suggests whoever plants a walnut tree will die as soon as the tree can "see" the sea. Most planting is done by field rats (subfamily Murinae). In Flanders, a folk saying states: "By the time the tree is big, the planter surely will be dead." (Dutch: Boompje groot, plantertje dood). These sayings refer to the relatively slow growth rate and late fruiting of the tree.

Benevento in southern Italy is the home of an ancient tradition of stregoneria. The witches of Benevento were reputed to come from all over Italy to gather for their sabbats under the sacred walnut tree of Benevento. In 1526, Judge Paolo Grillandi wrote of witches in Benevento who worship a goddess at the site of an old walnut tree.[23] This legend inspired many cultural works, including the 1812 ballet Il Noce di Benevento (the walnut tree of Benevento) by Salvatore Viganò and Franz Xaver Süssmayr, a theme from which was adapted into a violin piece called Le Streghe by Niccolò Paganini. The Beneventan liqueur Strega depicts on its label the famous walnut tree with the witches dancing under it.

See also

References

  1. ^ a b Rivers, M.C.; Allen, D.J. (2017). "Juglans regia". The IUCN Red List of Threatened Species. 2017: T63495A61526700. doi:10.2305/IUCN.UK.2017-3.RLTS.T63495A61526700.en.
  2. ^ "Juglans regia". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA). Retrieved 2019-07-06.
  3. ^ "Walnut". Online Etymology Dictionary. 2021.
  4. ^ L.C. van Zyl (2009). "Grafting of Walnut (Juglans regia L.) with Hot Callusing Techniques Under South African Conditions" (PDF). University of the Free State. Archived from the original (PDF) on 2012-09-07. Retrieved 2011-03-06.
  5. ^ a b "Walnuts: Australia - Nuts". Austnuts.com.au. Archived from the original on 2010-11-29. Retrieved 2012-08-23.
  6. ^ D.S. Hill, Skegness, Lincs, United Kingdom: Pests of Crops in Warmer Climates and Their Control p.651, Springer Science+Business Media, 2008
  7. ^ "Ornamental Tree Photography - NZ Plant Pics Photography ornamental garden trees". Nzplantpics.com. Archived from the original on 2013-03-02. Retrieved 2012-08-23.
  8. ^ "English walnuts - profile". Agricultural Marketing Resource Center. Archived from the original on September 28, 2006.
  9. ^ Walter Fox Allen. "How to grow English walnuts". WalnutsWeb. Archived from the original on April 25, 2009.
  10. ^ Juglans Regia (in Spanish)
  11. ^ ResearchGate (November 2014). "Nutritive Value of Persian Walnut (Juglans regia L.) Orchards". ResearchGate.
  12. ^ Beer, Ruth; Kaiser, Franziska; Schmidt, Kaspar; Ammann, Brigitta; Carraro, Gabriele; Grisa, Ennio; Tinner, Willy (2008-03-01). "Vegetation history of the walnut forests in Kyrgyzstan (Central Asia): natural or anthropogenic origin?". Quaternary Science Reviews. 27 (5): 621–632. Bibcode:2008QSRv...27..621B. doi:10.1016/j.quascirev.2007.11.012. ISSN 0277-3791.
  13. ^ Hemery 1998
  14. ^ Theophrastus, Enquiry into Plants III.6.2, III.14.4
  15. ^ "FAO corporate document repository: Walnut".
  16. ^ "Production of shelled walnuts in 2019, Crops/Regions/World list/Production Quantity (pick lists)". UN Food and Agriculture Organization, Corporate Statistical Database (FAOSTAT). 2020. Retrieved 23 January 2021.
  17. ^ "Walnuts in California". Fruit and Nut Research and Information Center, University of California at Davis. 2021. Retrieved 23 January 2021.
  18. ^ Boyce, A.M. (December 1929). "The Walnut Husk Fly (Rhagoletis juglandis Cresson)". Journal of Economic Entomology. 22: 861–866.
  19. ^ "Allergy information for walnut (Juglans regia)". Manchester Academic Health Science Centre, University of Manchester. 18 October 2006. Retrieved 22 January 2021.
  20. ^ Teuber, Suzanne S.; Jarvis, Koren C.; Dandekar, Abhaya M.; Peterson, W. Rich; Ansari, Aftab A. (1999). "Identification and cloning of a complementary DNA encoding a vicilin-like proprotein, Jug r 2, from English walnut kernel (Juglans regia), a major food allergen". Journal of Allergy and Clinical Immunology. 104 (6): 1311–1320. doi:10.1016/S0091-6749(99)70029-1. PMID 10589017.
  21. ^ Ozkhan, Gulcan; Koyuncu, M. Ali (2005). "Physical and chemical composition of some walnut (Juglans regia L.) genotypes grown in Turkey" (free). Grasas y Aceites. 56 (2): 141–146. doi:10.3989/gya.2005.v56.i2.122.
  22. ^ Elmore, Francis H., 1944, Ethnobotany of the Navajo, Santa Fe, NM. School of American Research, page 39
  23. ^ Grimassi, Raven. Encyclopedia of Wicca & Witchcraft. Llewellyn Worldwide (2000). p. 454.

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Juglans regia: Brief Summary

provided by wikipedia EN

Juglans regia, the Persian walnut, English walnut, Carpathian walnut, Madeira walnut, or especially in Great Britain, common walnut, is an Old World walnut tree species native to the region stretching from the Balkans eastward to the Himalayas and southwest China. It is widely cultivated across Europe and the Caucasus.

It is the origin of cultivated varieties which produce the edible walnut, consumed around the world. China is the major commercial producer of walnuts.

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