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Lifespan, longevity, and ageing

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Maximum longevity: 13.7 years (wild)
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Distribution

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Marbled godwits are found on the shores of the western United States, from the the Gulf of Mexico all the way up to Alaska in the winter months. They migrate to the northern plains of the U.S. and into the boreal forests of Canada in the summer. Isolated populations also breed in Alaska and in southwestern James Bay in Canada.

Biogeographic Regions: nearctic (Native )

Other Geographic Terms: holarctic

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Associations

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Marbled godwits were targeted by hunters in the 1800’s because they lived in huge flocks which were easy to hunt, and because they are large birds with tasty meat. Today, they are preyed upon by raccoons (Procyon lotor) and skunks (Mephitis) when nesting close to developed areas.

Known Predators:

  • raccoons (Procyon lotor)
  • skunks (Mephitis)
  • humans (Homo sapiens)
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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Morphology

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Marbled godwits are large tawny brown shorebirds with long legs and slightly upturned bills. They have tawny buff feathers which appear darker on top than underneath. Their underwings are mottled brown with cinnamon and are distinctive in flight. In winter, they are plain underneath, but during the breeding season they have dark barring on their breasts and bellies. Legs are gray or blue-gray and beaks are bright pink to orange. Marbled godwits are most similar to bar-tailed godwits (Limosa lapponica), which are slightly smaller with shorter legs and lack the bright cinnamon underwing of marbled godwits. They range from 42 to 48 cm and weigh 285 to 454 g. Females are larger than males. Their wingspan ranges from 74 to 78 cm.

Range mass: 285 to 454 g.

Range length: 42 to 48 cm.

Range wingspan: 74 to 78 cm.

Sexual Dimorphism: female larger

Other Physical Features: endothermic ; bilateral symmetry

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Life Expectancy

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Marbled godwits have a long lifespan. The record for the oldest in the wild was 30 years old.

Range lifespan
Status: wild:
20 to 30 years.

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Habitat

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Marbled godwits are the most widespread godwit species. They breed in grasslands or wetlands in the northern prairies of the United States and Canada, especially those lacking in dense or tall vegetation. They are also found in temporary ponds, as well as pastures and hay fields. More northern populations are found in wet tundra or open taiga, lowland meadows and bogs, and in coastal marshes. During migration and winter, they are found in wetlands and marshes, shallow ponds, coastal estuaries, mudflats, salt marshes, and sandy beaches.

Range elevation: 0 to 100 m.

Habitat Regions: saltwater or marine ; freshwater

Terrestrial Biomes: tundra ; taiga ; savanna or grassland ; forest

Aquatic Biomes: lakes and ponds; temporary pools; coastal

Wetlands: marsh ; bog

Other Habitat Features: agricultural ; estuarine

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Trophic Strategy

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The diet of marbled godwits varies with the season and location. In winter or on the coast, primary foods include annelid worms (Polychaeta), small bivalves (Bivalvia), crabs, and earthworms (Lumbricina). In summer or while inland, primary foods include insects such as grasshoppers (Orthoptera), aquatic plant tubers, leeches, and small fish. Marbled godwits move slowly while feeding, probing for food underneath the mud with their sensitive bill. They often insert their entire bill into the mud, and their head remains totally submerged at times.

Animal Foods: fish; insects; mollusks; terrestrial worms; aquatic or marine worms; aquatic crustaceans

Plant Foods: roots and tubers; seeds, grains, and nuts

Primary Diet: carnivore (Insectivore , Molluscivore , Vermivore)

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Associations

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Marbled godwits are commonly seen in social flocks with whimbrels (Numenius phaeopus) and long-billed curlews (N. americanus) along shorelines searching for food. They serve as host to several parasites such as avian botulism, caused by the bacterium Clostridium botulineum. Endoparasites include species-specific nematodes (Skrjabinoclava kristscheri), and other nematodes common in shorebirds (Ancyracanthopsis winegardi). Other nematodes infecting them include Sobolevicephalus lichtenfelsi, Viktoracana limosae, V. capillaris, and Stellocaronema skrjabiniin. Ectoparasites affecting them include mites such as Austromenopon limosae, Actornithophilus limosae, Carduiceps clayae, Lunaceps clayae, Rotundiceps cordatus, Saemundssonia.

Mutualist Species:

  • whimbrels (Numenius phaeopus)
  • long-billed curlews (N. americanus)

Commensal/Parasitic Species:

  • nematodes (Skrjabinoclava kristscheri)
  • nematodes (Ancyracanthopsis winegardi)
  • nematodes (Sobolevicephalus lichtenfelsi)
  • nematodes (Viktoracana limosae)
  • nematodes (Viktoracana capillaris)
  • nematodes (Stellocaronema skrjabiniin)
  • mites (Austromenopon limosae)
  • mites (Actornithophilus limosae)
  • mites (Carduiceps clayae)
  • mites (Lunaceps clayae)
  • mites (Rotundiceps cordatus)
  • mites (Saemundssonia)
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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Benefits

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Birding groups and tourism are attracted to marbled godwits. They even have days named after them called the Godwit Days. This is a time for lectures, field trips, boat excursions and workshops all based on marbled godwits.

Positive Impacts: food ; ecotourism

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Benefits

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There are no known negative economic impacts caused by marbled godwits.

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Conservation Status

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Marbled godwits are a species of least concern according to the IUCN Red List, though they are protected under the United States Migratory Bird Act. The Canadian Wildlife Service estimates the population at 171,500 birds. They were common in the 1800s, but were over-hunted in the early 1900s. Protection from hunting has helped the population rebound, but the destruction of grassland breeding habitat now limits the population. Marbled godwits require wetlands for breeding. As wetland ecosystems decline, marbled godwits will begin to migrate to alternate breeding grounds. After they migrate, their original breeding grounds are more difficult to protect as they no longer live in those areas.

US Migratory Bird Act: protected

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Catherine Kent, Special Projects
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Behavior

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Marbled godwits communicate by calling and physical displays, which are used especially for mate selection and interaction with predators. The call of marbled godwits is nasally, and is described as a slightly crowing or laughing "ah, ha" or "ahk." A specific kind of call is uttered when the individual calling enters a group, and is believed to decrease aggression against the newcomer.

Communication Channels: acoustic

Other Communication Modes: choruses

Perception Channels: visual ; tactile ; acoustic ; chemical

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Robert Sorensen, Minnesota State University, Mankato
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Reproduction

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Marbled godwits arrive at their breeding range in April or May in more southern populations, and form monogamous pairs. To attract a female, the male performs a high, circling flight display, followed by a steep dive. The male selects a nest site in a dry spot with short grass and starts a shallow scrape. If the female approves it, both will add grass, and sometimes a canopy of grass is then arched over the nest.

Mating System: monogamous ; cooperative breeder

Marbled godwits breed once per year between May and August. They form loose colonies without obvious territorial boundaries. Nests are constructed on the ground by males and approved by females, usually in an area with short vegetation. Nests are shallow depressions in the ground, lined with grasses and lichen or moss and leaves.

Females almost always lay 4 eggs, and rarely lay 3 or 5. Eggs are pale buff or olive with dark brown or purplish-gray spots or blotches. Both parents incubate the eggs, which are believed to hatch in 24 to 26 days. Parents sometimes join together to mob potential predators to defend young. At birth, precocial young are covered in down, have open eyes, and can walk and attempt to feed at birth. Young marbled godwits leave the nest after 1 to 2 days, and chicks fledge in 26 to 30 days after hatching.

Breeding interval: Marbled godwits breed once per year.

Breeding season: Marbled godwits breed from May to August.

Range eggs per season: 3 to 5.

Range time to hatching: 24 to 26 days.

Range time to independence: 15 to 26 days.

Range age at sexual or reproductive maturity (female): unknown (low) years.

Range age at sexual or reproductive maturity (male): unknown (low) years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

Both parents incubate the eggs for 24 to 26 days. They protect and tend the young for the first 15 to 26 days, after which the female usually leaves. The male stays with the young until they can fly.

Parental Investment: male parental care ; female parental care ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Male, Female); pre-weaning/fledging (Protecting: Male, Female); pre-independence (Protecting: Male)

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Kennedy, B. 2013. "Limosa fedoa" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Limosa_fedoa.html
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Brooks Kennedy, Minnesota State University, Mankato
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Marbled godwit

provided by wikipedia EN

The marbled godwit (Limosa fedoa) is a large migratory shorebird in the family Scolopacidae. On average, it is the largest of the four species of godwit.

Taxonomy

In 1750 the English naturalist George Edwards included an illustration and a description of the marbled godwit in the third volume of his A Natural History of Uncommon Birds. He used the English name "The Greater American Godwit". Edwards based his hand-coloured etching on a preserved specimen that had been brought to London from the Hudson Bay area of Canada by James Isham.[2] When in 1758 the Swedish naturalist Carl Linnaeus updated his Systema Naturae for the tenth edition, he placed the marbled godwit with godwits and ibises in the genus Scolopax. Linnaeus included a brief description, coined the binomial name Scolopax fedoa and cited Edwards' work.[3] The marbled godwit is now placed in the genus Limosa that was introduced in 1760 by the French zoologist Mathurin Jacques Brisson.[4][5] The genus name Limosa is from Latin and means "muddy", from limus, "mud". The specific epithet fedoa may be an Old English name for a godwit.[6] The word was mentioned by the English naturalist William Turner in 1544.[7][8]

Two subspecies are recognised:[5]

  • L. f. beringiae Gibson & Kessel, 1989 – breeds in Alaska and winters in the west United States. Averages shorter legs and bill than nominate subspecies.
  • L. f. fedoa (Linnaeus, 1758) – breeds in central, south-central Canada, and the north-central United States, winters in southern United States to northwest South America. A disjunct breeding population breeds in James Bay.

Description

The total length is 40–50 cm (16–20 in), including a large bill of 8–13 cm (3.1–5.1 in), and wingspan is 70–88 cm (28–35 in).[9] Body mass can vary from 240 to 520 g (8.5 to 18.3 oz). The average weight of 40 males was 326 g (11.5 oz) and that of 45 females was 391 g (13.8 oz). Bill length is from 73.9 to 131 mm (2.91 to 5.16 in). Among all the members of the sandpiper family, only the curlews attain sizes that significantly exceed this species.[10][11]

Adults have long blue-grey legs and a very long pink bill with a slight upward curve and dark at the tip. The long neck, breast and belly are pale brown with dark bars on the breast and flanks. The back is mottled and dark. They show cinnamon wing linings in flight.

Distribution and habitat

Marbled godwits breed in three distinct areas with their own unique route. The vast majority occur in mid-continental North America, followed by eastern Canada and the Alaska Peninsula, USA. In addition, the largest winter ranges are the Atlantic, Pacific and Gulf coasts of the US and Mexico.[12]

Godwits breeding in the western USA and Canada follow a route through the Utah stopover site, with a final arrival in the winter sites of Mexico and the Caribbean. Species breeding in eastern Canada migrate across the US, and stopover at sites along the Gulf of California and Mexico. Furthermore, those breeding in North and South Dakota winter in coastal Georgia.[12] The Bear River Migratory Bird Refuge located at Great Salt Lake in Utah (USA), is one of the most popular stopover sites for godwits in the spring and fall.

Behavior and ecology

Breeding

They nest on the ground, usually in short grass.

Food and feeding

These birds forage by probing on mudflats, in marshes, or at the beach (see picture below). When the tide is out, they eat. In short grass, they may pick up insects by sight. They mainly eat insects and crustaceans, but also eat parts of aquatic plants.

When the tide is in, they roost. They often sleep by standing on one leg and tucking their bill into their body (see picture below).[13]

Conservation

Their numbers were reduced by hunting at the end of the 19th century. Although they had recovered somewhat since that time, their population has declined in recent times as suitable habitat is used for farming.

Gallery

References

  1. ^ BirdLife International (2016). "Limosa fedoa". IUCN Red List of Threatened Species. 2016: e.T22693162A93387912. doi:10.2305/IUCN.UK.2016-3.RLTS.T22693162A93387912.en. Retrieved 12 November 2021.
  2. ^ Edwards, George (1750). A Natural History of Uncommon Birds. Vol. Part III. London: Printed for the author at the College of Physicians. p. 137, Plate 137.
  3. ^ Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 146.
  4. ^ Brisson, Mathurin Jacques (1760). Ornithologie, ou, Méthode Contenant la Divisio Oiseaux en Ordres, Sections, Genres, Especes & leurs Variétés (in French and Latin). Paris: Jean-Baptiste Bauche. Vol. 1, p. 48, Vol. 5, p. 261.
  5. ^ a b Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Sandpipers, snipes, coursers". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 14 October 2021.
  6. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 158, 227. ISBN 978-1-4081-2501-4.
  7. ^ Newton, Alfred (1893–1896). "Fedoa". A Dictionary of Birds. London: Adam and Charles Black. p. 248.
  8. ^ Turner, William (1903) [1544]. Turner on Birds: a short and succinct history of the principal birds noticed by Pliny and Aristotle first published by Doctor William Turner, 1544 (in Latin and English). Translated by Evans, A.H. Cambridge: Cambridge University Press. p. 45.
  9. ^ [1] (2011).
  10. ^ CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), ISBN 978-0-8493-4258-5.
  11. ^ Gratto-Trevor, C. L. (2020). Marbled Godwit (Limosa fedoa), version 1.0. In Birds of the World (A. F. Poole and F. B. Gill, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA.
  12. ^ a b Olson, Bridget E.; Sullivan, Kimberley A.; Farmer, Adrian H. (May 2014). "Marbled Godwit migration characterized with satellite telemetry". The Condor. 116 (2): 185–194. doi:10.1650/CONDOR-13-024.1. JSTOR 90008440. S2CID 86113359.
  13. ^ Lentz, Joan Easton (November 2005). Introduction to Birds of the Southern California Coast. University of California Press. p. 151. ISBN 9780520243217. Retrieved 27 December 2017.

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Marbled godwit: Brief Summary

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The marbled godwit (Limosa fedoa) is a large migratory shorebird in the family Scolopacidae. On average, it is the largest of the four species of godwit.

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Distribution

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North America

Reference

North-West Atlantic Ocean species (NWARMS)

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