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Asker, Akershus, Norge
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Figure 4.Habitats of the Antiperyphanes Complex. A River shore, Argentina: Neuquén: Rio Collón Curá, about 13 km S La Rinconada, 625m. On the sandy bank in the foreground Bembidion philippii is abundant, as is Bembidion (Nothonepha) sp. nr. lonae. Also occurring on the sand banks are Bembidium orregoi and Bembidion (Nothonepha) eburneonigrum. On the upper sand banks across the river are Bembidium mandibulare, and in the gravel are Bembidium spinolai B Edges of snowfields at Chile: Reg. IX: Volcán Lonquimay, 1910m. Habitat of Bembidion (Notholopha) sexfoveolatum C Open high-elevation grassland at Ecuador: Pichincha: Paso de la Virgen, 4060m, habitat of three species of subgenus Ecuadion: Bembidion chimborazonum, Bembidion guamani, and Bembidion humboldti D Leaf litter in cloud forest, Ecuador: Pichincha: Reserva Yanacocha, 0.1152°S, 78.5837°W, 3540m, habitat of Bembidion andersoni, Bembidion georgeballi, and Bembidion onorei.
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Asker, Akershus, Norge
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Figure 5.Adults of subgenus Nothonepha. A Bembidion tetrapholeon (black form), Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100781 B Bembidion tetrapholeon (orange form), Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100780 C Bembidion germainianum, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100782 D Bembidion lonae, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100786 E Bembidion eburneonigrum, Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785 F Bembidion tucumanum, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100783 G Bembidion engelhardti engelhardti, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100784. Scale bar 1 mm.
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Asker, Akershus, Norge
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Figure 5.Adults of subgenus Nothonepha. A Bembidion tetrapholeon (black form), Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100781 B Bembidion tetrapholeon (orange form), Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100780 C Bembidion germainianum, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100782 D Bembidion lonae, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100786 E Bembidion eburneonigrum, Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785 F Bembidion tucumanum, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100783 G Bembidion engelhardti engelhardti, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100784. Scale bar 1 mm.
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Asker, Akershus, Norge
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Figure 14.Inner surface of the anterior portion of the abdomen, soft tissue removed. The abdomen is slightly tilted to the left. sp: sclerotized patch on membrane that bounds the front of the abdomen on the ventral side. All species shown are members of the Bembidion Series; A–F are members of the Antiperyphanes Complex; A, C and D are members of the subgenus Nothonepha A Bembidion tetrapholeon, DRM voucher V100766 B Bembidium mandibulare, DRM voucher V100805 C Bembidion sp. nr. lonae, DRM voucher V100803 D Bembidion lonae, DRM voucher V100802 E Bembidium philippii, DRM voucher V100804 F Bembidion chimborazonum, DRM voucher V100809 G Bembidion charile Bates, DRM voucher DNA1171 H Bembidion quadrimaculatum oppositum Say, DRM voucher V100807. Scale bar 0.1 mm.
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Tjærbæk Toholt Skov
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Figure 7.Tree of highest likelihood found for the combined, seven-gene matrix with partitioning scheme chosen by PartitionFinder. Only members of the Bembidion Series shown; more distant outgroups not depicted. Scale bar: 0.01, branch lengths as reconstructed by RAxML.
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Tjærbæk Toholt Skov
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Figure 8.Maximum likelihood bootstrap tree showing only those clades appearing in 90% of the bootstrap replicates; taxa outside of the Antiperyphanes Complex not shown. Numbers below branches indicate maximum likelihood bootstrap percentage / parsimony bootstrap percentage. Circled letters on branches correspond to groups documented in Table 3.
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Tjærbæk Toholt Skov
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Figure 9.Left lateral region of the prothorax and mesothorax; top of each photograph is anterior. msst: mesosternum; msepst: mesepisternum; msep: mesepimeron (note that the boundaries between these sclerites are not evident externally in A. A Bembidion (Nothonepha) tetrapholeon, DRM voucher V100810 B Bembidion (Antiperyphanes) zanettii, DRM voucher V100811. Scale bar 0.1 mm.
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Tjærbæk Toholt Skov
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Figure 15.Pronotum of Bembidion tetrapholeon, DRM voucher V100781. Scale bar 0.1 mm.
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Asker, Akershus, Norge
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Figure 16.Humeral region of left elytron and posterior corner of pronotum of three Bembidion (Nothonepha) species. A Bembidion tetrapholeon, DRM voucher V100781 B Bembidion lonae, DRM voucher V100786 C Bembidion germainianum, DRM voucher V100782. Arrows show the anterior end of the lateral elytral groove and bead. Scale bar 0.1 mm.
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Rugård Sønderskov
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Figure 17.Male aedeagus of Bembidion tetrapholeon. A Black form, Chile: Region X, Rio Pullinque at Puente Huanehue, 8 km NE Panguipulli, DRM voucher DNA1752 B Black form, Chile: Reg. X, Chiloé: Rio Puntra at route 5, DRM voucher DNA2236 C Orange-spotted form, Argentina: Neuquén: Rio Pichi Traful nr Lago Traful, DRM voucher DNA2564 D Orange-spotted form, Argentina: Neuquén: Rio Pichi Traful nr Lago Traful, DRM voucher DNA2555. Scale bar 0.1 mm.
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Rugård Sønderskov
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Figure 18.Geographic distribution of Bembidion tetrapholeon. Circles filled in black are based upon specimens I have examined and that have been sequenced; circles filled in gray are based upon specimens identified by Luca Toledano. Five cities are included as landmarks.
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Rugård Sønderskov
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Figure 6.Habitats of the subgenus Nothonepha. A Habitat of Bembidion (Nothonepha) eburneonigrum (on sand patches) and Bembidion (Nothonepha) sp. nr. lonae (on sand patches and in gravel). This habitat (Chile: Reg. IX: Rio Allipén at route 119, 132m) is also home to Bembidion spinolai Solier and Bembidion rufoplagiatum B Habitat of Bembidion (Nothonepha) tucumanum, Bembidion (Nothonepha) lonae, and Bembidion (Nothonepha) engelhardti (Argentina: Mendoza: Salinas del Diamante, 1280m). The beetles are common under rocks and around vegetation on the salt-encrusted clay and sand banks of this saline pond; in the same habitat Bembidion (Notaphus) cillenoides Jensen-Haarup and two other Notaphus are common C On the sand shores of this desert river Bembidion (Nothonepha) germainianum, Bembidion (Nothonepha) engelhardti, and Bembidion (Nothonepha) lonae are common (Argentina: Neuquén: Rio Salado at route 40, 725m) D Type locality of Bembidion (Nothonepha) tetrapholeon (Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m). The beetles are found under rocks along the river shore; Bembidion rufoplagiatum is also common in this habitat.