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Canoe Cedar

Thuja plicata Donn ex D. Don

Associations

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In Great Britain and/or Ireland:
Foodplant / pathogen
Armillaria mellea s.l. infects and damages Thuja plicata

Foodplant / saprobe
immersed, opening by little lids apothecium of Didymascella thujina is saprobic on dead, attached leaf of Thuja plicata
Remarks: season: 6-8

Foodplant / saprobe
immersed acervulus of Truncatella coelomycetous anamorph of Truncatella hartigii is saprobic on bark of Thuja plicata

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Comments

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The range of Thuja plicata consists of a Coast Range--Cascade Range segment from southeastern Alaska to northwestern California (between 56° 30' and 40° 30' N latitude) and a Rocky Mountains segment from British Columbia to Idaho and Montana (between 54° 30' and 45° 50' N latitude).

Thuja plicata is an important timber tree. Its soft but extremely durable wood is valued for home construction, production of shakes and shingles, and many other uses. Native Americans of the Northwest Coast used it to build lodges, totem poles, and seagoing canoes. Many cultivars are grown for ornament, and the species is managed for timber in Europe and New Zealand.

Western redcedar ( Thuja plicata ) is the provincial tree of British Columbia.

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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Description

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Trees to 50(-75) m tall; trunk to 2(-5) m d.b.h., often buttressed at base; bark reddish brown or grayish brown, fibrous, fissured; crown conical; branches arching; branchlets pendulous. Leaves on upper side of branchlets glossy green, (1-)3-6 mm, apex acuminate; lateral leaves longer than facial leaves, apex straight. Pollen cones reddish, 1-3 mm. Seed cones brown, ellipsoid, 1-1.4 cm; fertile cone scales 4-6. Seeds reddish-brown, 4-7.5 mm (including wings).
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 63 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Description

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Trees to 50(--75) m, sometimes stunted in harsh environments; trunk to 2(--5) m diam., often buttressed at base; crown conical. Bark reddish brown or grayish brown, 10--25 mm thick, fibrous, fissured. Branches arching, branchlets pendent. Leaves of branchlets (1--)3--6 mm (sprays sometimes bearing only very small leaves), apex acute, with white markings on abaxial surface when fresh, glossy green on adaxial surface of branchlets. Pollen cones 1--3 mm, reddish. Seed cones ellipsoid, 10--14 mm, brown; fertile scales 2--3 pairs, each with evident, nearly terminal, deltate projection. Seeds 8--14 per cone, 4--7.5 mm (including wings), reddish brown. 2 n = 22.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of North America @ eFloras.org
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Flora of North America Editorial Committee
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eFloras.org
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Distribution

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Alta., B.C.; Alaska, Calif., Idaho, Mont., Oreg., Wash.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
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Flora of North America Editorial Committee
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eFloras.org
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Habitat

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On various substrates, commonly in moist sites, mixed coniferous forests, usually not in pure stands; 0--1500(--2000)m.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
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Flora of North America Editorial Committee
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Habitat & Distribution

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Cultivated. Jiangsu, Jiangxi [native to W Canada, NW United States]
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 63 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Synonym

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Thuja gigantea Nuttall.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 4: 63 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
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eFloras.org
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eFloras

Medicinal Uses by Native Americans of the US and Canada

provided by EOL authors

Thuja plicata, or Western Red Cedar, is a cultural keystone species for many Native American people of the Pacific North West of the United States and Canada; T. plicata is so important and provides so many products that native peoples of this region are often called "People of the Cedar", and the Kakawaka'wakw in particular call it "the tree of life" (Garibaldi and Turner 2004; Gunther 1945). Many Native American groups use it for crafting implements, structures, clothing, and ceremonial items as well as medicinally (Gunther 1945).

The Bella Coola of British Colombia treat a variety of symptoms with three common preparations of Thuja plicata: leaf decoctions, infusions of leaves, and, poultices of pounded bough tips and eulachon (candlefish) grease. Decoctions (and compound decoctions of powdered leaves) are used externally for internal pains such as stomach pain. While leaf infusions, and, pounded bough tip poultices applied, are both used externally for rheumatism, heart trouble and neck swelling. The two differ in that leaf infusions are also used for coughs, while the poultices can also be used for bronchitis and stomach pain. Soft bark is used like a bandage to cover wounds and skin applications (Moerman 1998).

In additional to the Bella Coola, many other groups overlap in the use of this plant to treat coughs and respiratory issues. The Makah and Nez Perce both use bough infusion, while the Skagit use leaf decoctions. Nez Perce also treat colds with bough infusions, which Cowlitz treat with decoctions of plant tips and roots. The Klallam (Clallam), natives of Olympic Peninsula , Washington and the southern shore of Vancouver Island Columbia, use decoctions of small branches for tuberculosis(Moerman 1998; Gunther 1945).

Other common uses of the plant are as a skin application, and as a solution to diarrea. Moxa (a dried herb substance burned on or near the skin) of the inner bark is used as a counter irritant for skin by the Haisla while the Kwakiutl use inner bark to make a poultice for application to carbuncles and use shredded bark to cauterize sores and swellings the of feet. The Colville use bough infusions of it as a solution for dandruff and scalp issues and for soaking arthritic and rheumatic joints, weak infusions are also taken for arthritis and rheumatism as the Bella Coola do. The Hanaksiala and Nez Perce both use leaves as antidiarrheals (Moerman 1998).

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Amy Chang
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Amy Chang
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Broad-scale Impacts of Plant Response to Fire

provided by Fire Effects Information System Plants
More info for the term: prescribed fire

Hamilton's Research Papers (Hamilton 2006a, Hamilton 2006b) provide
information on prescribed fire and postfire response of plant
community species, including western redcedar, that was not available when
this species review was originally written.
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Common Names

provided by Fire Effects Information System Plants
western redcedar
western red-cedar
Pacific redcedar
giant redcedar
arborvitae
canoe-cedar
shinglewood
giant arborvitae
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Description

provided by Fire Effects Information System Plants
More info for the terms: duff, tree

Western redcedar is a large, native, long-lived, evergreen tree
[8,50,57,80].  At maturity it is generally 70 to 100 feet (21-30 m)
tall, sometimes 130 feet (40 m), with a tapering trunk 2 to 4 feet
(0.6-1.2 m) in diameter, sometimes 6 feet (1.8 m) or more.  On some
sites west of the Cascades, old-growth western redcedar often attains
basal diameters of 8 to 10 feet (2.4-3 m) and heights of 200 feet (61
m).  The largest known western redcedars are believed to be 1,000 years
old or more [8].

Western redcedar has a swollen or buttressed base, pointed conical
crown, and horizontal branches curving upward at the tips [80].  The
leaves are scalelike, flattened and 0.05 to 0.1 inches (1.5-3 mm) long.
The twigs are flattened, in fanlike sprays and slightly drooping.  The
bark is thin, fibrous and stringy or shreddy.  Thickness varies from 0.5
to 1 inch (1.3-2.5 cm) [51].  The cones are
clustered near the ends of
twigs and become turned up on short stalks [80].  Western redcedar
retains its lower limbs except when in densely crowded stands [8].

Western redcedar roots are extensive.  Tap roots are poorly defined or
nonexistent, but fine roots develop a profuse, dense network.  Root
systems tend to be shallower and less extensive on wet soils than on
deep, moderately dry soils.  When a thick duff layer is present, many
western redcedar roots lie in the duff rather than in the underlying
soil [50,51].
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Distribution

provided by Fire Effects Information System Plants
More info for the term: forest

Western redcedar occurs along the Pacific Coast from the southern part
of the Alaska Panhandle through British Columbia, western Washington,
and western Oregon, reaching into the coastal redwood forest of northern
California [8,50,54,57].  Inland from the coast it occupies a contiguous
band east of the Cascade Range from central Oregon to southern British
Columbia [54].  Much farther inland a disjunct population occurs along
the west slopes of the Rocky Mountains from Prince George, British
Columbia, to northeastern Washington, northern Idaho, and western
Montana [50,54,57].
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cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Ecology

provided by Fire Effects Information System Plants
More info for the terms: fire interval, fire regime, frequency, mean fire interval

Western redcedar fire resistance is low to moderate [21].  Its thin
bark, shallow root system, low dense branching habit, and highly
flammable foliage make it susceptible to fire damage [21,59].  However,
it often survives fire because of it large size [21].  Old western
redcedar trees are commonly fire scarred in northern Idaho [51].
Western redcedar is more severely damaged by fire than any of its
associates along the coast region but is less susceptible than Engelmann
spruce (Picea engelmannii), western hemlock, and subalpine fir in
interior regions [50].

The frequency of fire in western redcedar stands tends to be low
[13,73].  In most of the western redcedar forests from southern British
Columbia to northern California moderate to severe wildfires occur at
long intervals between 50 to 350 years [10].  In streamside and seepage
areas dominated by western redcedar the mean fire interval is greater
than 200 years.  In western redcedar habitats on lower and middle slopes
the mean fire interval is 50 to 150 years [82].  In the
Selway-Bitterroot Wilderness those stands dominated by western redcedar
had the longest fire regime [29].

FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find FIRE REGIMES".
license
cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Management Considerations

provided by Fire Effects Information System Plants
More info for the terms: duff, fuel, tree

Riparian stringers supporting western redcedar may act as firebreaks
because the moist duff does not readily burn [21].  Old-age western
redcedar stands have heavy fuel loads, but a large proportion of this
material is in the form of deep duff layers and downed, rotting log
material.  These stands could support slow moving fires at best; once
ignited, however, such heavy fuel materials could support long-lasting
fires [29].
 
Fire-killed western redcedar often shows little deterioration even after
5 years.  The bark usually remains intact on dead tree for 5 years.
Fire mortality produces no immediate reduction in strength of western
redcedar poles, and some large trees remain salvageable for almost 100
years after being killed by fire [51].

When slash from decadent western redcedar-western hemlock stands was
burned, a greater proportion of western redcedar than of western hemlock
slash was consumed.  This was a result of greater longitudinal and
horizontal fracturing of the western redcedar.  When fracturing does not
occur, western hemlock slash is at least as flammable as western
redcedar slash.  Fire spreads faster in western redcedar when the slash
from both species is 1 year old.  Western redcedar slash does not drop
its foliage.  The slash of western redcedar is less flammable when
chipped.  One study showed that the fire hazard normally associated with
cutting of western redcedar poles was reduced by skidding entire
pole-size trees to the landing, where the slash was chipped and blown
over the edge [51].

Slash from western hemlock-western redcedar-Alaska-cedar forests produce
greater nutrient losses to the atmosphere when the slash composition has
a greater proportion of Alaska-cedar and western redcedar.  One can
expect smaller nutrient losses when western hemlock makes up the
majority of the slash [23].  For further details on slash burning of
western redcedar refer to the fire case study in the Alaska-cedar Fire
Effects Information System species review.
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Growth Form (according to Raunkiær Life-form classification)

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More info for the term: phanerophyte

  
   Phanerophyte
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat characteristics

provided by Fire Effects Information System Plants
More info for the terms: fern, herb, natural, organic soils, shrub

Western redcedar grows best in maritime climates with cool, cloudy
summers and wet, mild winters.  In drier areas west of the Cascades,
western redcedar becomes abundant only on wet sites such as ravines,
along streams, or on poorly drained bottomlands.  Near its range limits
in the drier mountains east of the Cascade crest, western redcedar grows
almost exclusively in narrow canyons, where its roots are irrigated all
summer by a mountain stream [8].  In Glacier National Park and the
Selway-Bitterroot Wilderness in Idaho and Montana, western redcedar is
dominant in wet ravines and poorly drained depressions [50].

Precipitation and temperature:  Western redcedar occurs on sites that
receive from 35 inches (890 mm) of annual precipitation to more than 260
inches (6,600 mm), mostly as winter rainfall [50].  Western redcedar is
not resistant to frost and is sometimes damaged by freezing temperatures
in late spring or early autumn.  When sufficient precipitation is
present, low temperatures appear to limit western redcedar's range.  The
northern limits of western redcedar lie between the 52 and 53 degree
Fahrenheit (11.1-11.7 deg C) mean summer temperature isotherms in
southeastern Alaska [50].  Bottomland frost pockets in northern Idaho
are commonly occupied by subalpine fir (Abies lasiocarpa) rather than
western redcedar [51].

Soils:  Western redcedar can tolerate a wide range of soil.  It is found
on all soil textures and parent materials on Vancouver Island.  Coarse
sandy soils are not well suited to the establishment and growth of
western redcedar in northern Idaho and northeast Washington, but rocky
slopes with limited soil development support western redcedars in
southeastern Alaska.  Poorly drained organic soils support redcedar
south of Petersburg, Alaska.  It grows well on shallow soils over chalk
and can tolerate both acid and alkaline soils conditions.  It is able to
survive and grow on soils that are low in nutrients and is found on such
soils over much of its natural range.  However, productivity may be
improved by fertilization [50].

Elevation:  Elevational ranges of western redcedar have been reported as
follows [19,50,51]:

        Alaska - 0 to 3,000 feet (0-910 m)
        British Columbia - 0 to 3,900 feet (0-1,190 m)
        Oregon - 0 to 7,500 feet (0-2,290 m)       
        northern Rocky Mountains - 2,000 to 5,900 feet (610-1,798 m)

In coastal regions, western redcedar is commonly associated with the
following shrub and herb species:  dwarf Oregon grape (Mahonia nervosa),
stink currant (Ribes bracteosum), Alaska blueberry (Vaccinium
alaskaense), box blueberry (V. ovatum), Pacific rhododendron
(Rhododendron macrophyllum), salal (Gaultheria shallon), threeleaf
anemone (Anemone deltoidea), deerfern (Blechnum spicant), slough sedge
(Carex obnupta), and evergreen violet (Viola sempervirens) [50,51].

In interior regions western redcedar is commonly associated with the
following shrub and herb species:  mountain alder (Alnus incana spp.
tenuifolia), Oregon grape (Mahonia repens), common juniper (Juniperus
communis), red raspberry (Rubus idaeus), blue huckleberry (Vaccinium
globulare), Rocky Mountain honeysuckle (Lonicera utahensis), gold thread
(Coptis occidentalis), roundleaf alumroot (Heuchera cylindrica), pine
drops (Pterospora andromedea), and green pyrola (Pyrola chlorantha)
[35,50,51,61].

Common shrub and herb associates of both coastal and interior regions
are as follows:  western serviceberry (Amelanchier alnifolia),
thimbleberry (Rubus parviflorus), oceanspray (Holodiscus discolor),
Devil's club, common snowberry (Symphoricarpos albus), lady fern,
western swordfern (Polystichum munitum), prince's-pine (Chimaphila
umbellata), bunchberry dogwood (Cornus canadensis), false Solomon's-seal
(Smilacina stellata), and Pacific trillium (Trillium ovatum) [50,51].
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Cover Types

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More info on this topic.

This species is known to occur in association with the following cover types (as classified by the Society of American Foresters):

   210  Interior Douglas-fir
   212  Western larch
   213  Grand fir
   215  Western white pine
   218  Lodgepole pine
   221  Red alder
   222  Black cottonwood - willow
   223  Sitka spruce
   224  Western hemlock
   225  Western hemlock - Sitka spruce
   226  Coastal true fir - hemlock
   227  Western redcedar - western hemlock
   228  Western redcedar
   229  Pacific Douglas-fir
   230  Douglas-fir - western hemlock
   231  Port-Orford-cedar
   232  Redwood
license
cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Ecosystem

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More info on this topic.

This species is known to occur in the following ecosystem types (as named by the U.S. Forest Service in their Forest and Range Ecosystem [FRES] Type classification):

   FRES20  Douglas-fir
   FRES21  Ponderosa pine
   FRES22  Western white pine
   FRES23  Fir - spruce
   FRES24  Hemlock - Sitka spruce
   FRES25  Larch
   FRES26  Lodgepole pine
   FRES27  Redwood
   FRES28  Western hardwoods
license
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bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Plant Associations

provided by Fire Effects Information System Plants
More info on this topic.

This species is known to occur in association with the following plant community types (as classified by Küchler 1964):

More info for the term: forest

   K001  Spruce - cedar - hemlock forest
   K002  Cedar - hemlock - Douglas-fir forest
   K003  Silver fir - Douglas-fir forest
   K004  Fir - hemlock forest
   K005  Mixed conifer forest
   K006  Redwood forest
   K012  Douglas-fir forest
   K013  Cedar - hemlock - pine forest
   K014  Grand fir - Douglas-fir forest
   K015  Western spruce - fir forest
license
cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Immediate Effect of Fire

provided by Fire Effects Information System Plants
More info for the term: duff

Western redcedar is commonly killed by fire.  Because of their large
size, however, old western redcedar trees can often survive if they are
not completely girdled by fire [21].  Shallow roots under the duff layer
are often scorched when the duff layer burns and even surface fires may
kill western redcedar [51].  Fire injury to roots can lead to fungal
infection, chronic stress, and growth losses [65].  The most common
causes of fire mortality are root charring and crown scorching [70].
license
cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Importance to Livestock and Wildlife

provided by Fire Effects Information System Plants
More info for the terms: cover, forest, tree

Black-tailed deer browse western redcedar seedlings and saplings all
year long in British Columbia, and Roosevelt elk feed on them during the
fall, winter, and spring.  Western redcedar constitutes one of the most
important conifer foods of black-tailed deer in the Coastal forest
region of southern Vancouver Island [51].  Western redcedar was more
severely browsed than Douglas-fir (Pseudotsuga menziesii), western
hemlock, or Pacific silver fir (Abies amabilis) on the Olympic
Peninsula.  Western redcedar is a major winter food for big game in the
northern Rocky Mountains [51].  An analysis of 69 stomach samples
collected from elk harvested along the Lochsa and lower Selway rivers
between January 1 and April 1 from 1960 through 1970 showed that western
redcedar leaves made up 5 percent of the total winter diet by weight
[72].  In western Washington, black bears remove western redcedar bark
and feed on the exposed sapwood [50].

Cattle browse western redcedar in preference to Douglas-fir in
northwestern Oregon, and sheep damaged western redcedar reproduction
more than that of other trees in northern Idaho [51].  Seeds of this
conifer were only occasionally taken by field mice in caged tests [52].

Old-growth stands of western redcedar provide hiding and thermal cover
for several wildlife species.  Bears, raccoons, skunks, and other
animals use cavities in western redcedar for dens [8].  In the southern
Selkirk Mountains of northern Idaho, northeastern Washington, and
adjacent British Columbia, grizzly bears have been known to use heavily
timbered western redcedar and western hemlock forests [44].  Western
redcedar is used as nest trees by cavity nesting bird species such as
yellow-bellied sapsuckers, hairy woodpeckers, tree swallows, chestnut
backed chickadees, and Vaux's swifts [45,49].
license
cc-publicdomain
bibliographic citation
Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Key Plant Community Associations

provided by Fire Effects Information System Plants
More info for the terms: association, codominant, forest, habitat type, series

Western redcedar commonly occurs as a dominant or codominant on
low-elevation moist sites.  In Montana, the western redcedar habitat
type series described by Pfister and others [60] occurs most extensively
in the Swan Valley and Mission Range, extends eastward locally to
Missoula, and forms small riparian stringers along major streams in the
Bitterroot Range west of Hamilton.  Western redcedar occurs as a
riparian dominance type on toe-slope seepages, moist benches, and wet
bottoms adjacent to streams [35].  Daubenmire and Daubenmire [16]
recognized three western redcedar communities in northern Idaho.
Western redcedar/pachistima (Pachistima myrsinites) is an upland
community, while western redcedar/Devil's club (Oplopanax horridus) and
western redcedar/ladyfern (Athyrium filix-femina) occur on bottomlands.
Western redcedar is sometimes found as a codominant with western hemlock
(Tsuga heterophylla) [16,35,60].  Published classifications identifying
western redcedar as a dominant or codominant are as follows:

Old-growth forests of the Canadian Rocky Mountains National Parks [3].
Preliminary plant associations of the southern Oregon Cascade Mountain
  Province [11].
Classification and management of riparian and wetland sites in
  northwestern Montana [13]. 
Forest habitat types of northern Idaho: a second approximation [15].
Forest Vegetation of eastern Washington and northern Idaho [16].
Fire ecology of Lolo National Forest habitat types [17].
Preliminary forest plant association management guide. Ketchikan area,
  Tongass National Forest [18].
Fire ecology of western Montana forest habitat types [21].
A guide to the interior cedar-hemlock zone, northwestern transitional
  subzone (ICHg), in the Prince Rupert Forest Region, British Columbia
  [33].
Riparian dominance types of Montana [35].
Classification and management of riparian sites in southwest Montana
  [36].
Soil classification as an aid to identifying forest habitat types in
  northern Idaho [54].
Forest habitat types of Montana [60].
Reference material Daubenmire habitat types [77].
Preliminary forest plant associations of the Stikine area, Tongass
  National Forest [78].
A study of the Vegetation of southeastern Washington and adjacent Idaho
  [81].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Life Form

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Tree
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Management considerations

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Insects and disease:  Western redcedar is a host for several
economically important insect species.  One of the most important is the
gall midge (Mayetiola thujae), which sometimes seriously damages western
redcedar seeds in Oregon, Washington, and British Columbia [50].
Seedlings are occasionally damaged by weevils (Steremnius carenatus) in
British Columbia, and large trees are killed by bark beetles
(Phloeosinus sequoiae) on poor sites in southeastern Alaska.  The
western redcedar borer (Trachykele blondeli) causes degradation
resulting in cull of sawtimber [50].

More than 200 fungi are found on western redcedar.  A leaf blight
(Didymascella thujina) infects second- and third-year nursery seedlings.
As much as 97 percent of the natural western redcedar regeneration may
be killed when this blight reaches epidemic proportions.  The most
important fungi attacking western redcedar are root butt and trunk rots.
Poria asiatiaa and P. albipellucida are the most important trunk rots
near the coast; P. asiatioa and Phellinus weiri are the most important
in the interior range.  Rots are most evident in old stands [50].

Animal damage:  Seedlings and saplings are often severely browsed by
deer, elk, and rodents.  Browse damage may be one of the most important
stand establishment problems [50].  Grazing by cattle in burned stands
in the western redcedar/queencup beadlily (Clintonia uniflora) habitat
type in southwestern Montana retards establishment of western redcedar
[36].

Other damaging agents:  Western redcedar is often windthrown in wet
environments, but it is windfirm on dry sites [50,51].  Western redcedar
is damaged by salt spray [50].  It is also sensitive to atmospheric
pollution.  Clay dust from a brick works in British Columbia produced a
columnar form in nearby western redcedars [51].

Silvicultural considerations:  Care must be exercised when logging sites
dominated by western redcedar due to the high water table.  Bottomland
sites should not be disturbed other than to salvage high-value trees or
to remove high-risk trees.  If harvested some dead and down logs should
be left to serve as a seedbed for western redcedar and western hemlock
regeneration.  Extensive disturbance of these sites could cause
irreparable damage [13].

Western redcedar should be grown in pure stands when saw-timber,
shingles, or shakes are the desired products.  Even-aged mixtures of
western redcedar and other conifers will be harvested either too early
for the western redcedar sawtimber or too late for the other conifers
when mixed-species, even-aged stands are clearcut.  Western redcedar can
be grown in mixed stands when poles are to be produced under even-aged
management regimes.  A nearly closed canopy should be maintained at all
times.  Open-grown western redcedar tend to develop poor form, excessive
limbs, and multiple tops [51].  Western redcedar is perhaps the most
valuable species for which uneven-aged systems are applicable in the
highly productive western redcedar and western hemlock habitat types of
the Inland West [26].

Response to release:  Because western redcedar is shade tolerant, it
should be treated to minimize shock from release through slow or timely
thinning treatments.  Western redcedar's ability to respond to release
varies with tree, stand, and site conditions.  An 80-year-old western
redcedar stand, with the overstory removed and thinned, responded with
increased growth rates up to 5 years after treatment.  However, 5 to 10
years after release, growth rates slowed, and root diseases became
apparent [26].  Releasing western redcedar saplings slowly over a 17
year period had good results.  The saplings responded favorably to
release with increased growth rates and a gradual increase in vigor.
Thinning western redcedar stands should occur prior to age 30.  Spacing
of 1 foot by 1 foot (0.3 by 0.3 m) is appropriate for most young stands.
This density provides good tree and stand development and retains the
options for future intermediate treatments [25,26].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Nutritional Value

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Relatively high concentrations of calcium and low concentrations of
nitrogen are nearly always present in western redcedar foliage.
Phosphorous concentrations are usually low [51].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Occurrence in North America

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     AK  CA  HI  ID  MT  OR  WA  AB  BC
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Other uses and values

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Perfumes, insecticides, medicinal preparations, veterinary soaps, shoe
polishes, and deodorants are made from western redcedar leaf oil.
Western redcedar extractives and residues are used in lead refining,
boiler-water additives, and glue extenders [50].  Western redcedar was
an extremely valuable tree to the Indians of the Northwest Coast,
providing materials for their shelters, clothing, dugout canoes, and
fishing nets [8,76].  Northwest Coast Indians shredded the inner layer
of bark so finely that it could be used for diapers and cradle padding
[8].

Western redcedar's drooping branches, thin fibrous bark, and flat sprays
of scalelike leaves make it an attractive ornamental.  When properly
trimmed western redcedar is an excellent hedge [8,41].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Phenology

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More info for the terms: cone, phenology, seed

The reproductive cycle of western redcedar occurs over approximately 16
months.  Phenology varies between coastal and interior regions.  For
trees in the middle of the coastal distribution on Vancouver Island,
pollen and seed cones develop in early June.  Pollen forms in late
February or early March of the second season.  Pollination occurs within
1 to 2 weeks usually in March but may begin as early as mid-February in
mild coastal areas or as late as early April at higher elevations.
Pollination in March is most common [57].  Fertilization occurs in late
May.  Cones mature in October.  West of the Cascade Range, cone maturity
is usually reached in 5 months, but in northern Idaho it takes 3 months.
Major seedfall occurs during October and November in both the interior
and coast range [50].  Dry warm weather can cause earlier seed release.
Some seeds may be retained in the cones and gradually shed throughout
the winter [19].  Where moisture and temperature conditions are
favorable, germination can occur in the autumn, winter, or spring [50].
Along the coast region, seeds generally germinate in either fall or
spring [8].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Plant Response to Fire

provided by Fire Effects Information System Plants
After fire, western redcedar will readily establish on bare mineral soil
seedbeds via off-site wind dispersed seeds [22,26].  Although unburned
soil benefits western redcedar regeneration more than soil that has been
scorched, slash burning favors western redcedar by creating more mineral
soil surfaces in cutover areas [50].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Post-fire Regeneration

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More info for the terms: root crown, secondary colonizer, tree

   Tree without adventitious-bud root crown
   Secondary colonizer - off-site seed
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Regeneration Processes

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More info for the terms: climax, competition, cone, duff, epigeal, layering, natural, seed, shrubs, stratification, tree

Seed production and dissemination:  Western redcedar reproduces from
seeds more readily in open, disturbed areas, such as clearcuts, than in
undisturbed stands [19].  Seed production normally begins when trees are
20 to 30 years old [73].  However, open-grown trees may produce seed by
age 10 [19,50].  Cones average about three to six seeds, but cones are
often numerous and heavy seed crops are common.  Average annual seed
crops vary from 100,000 to 1,000,000 seeds per acre
(247,000-2,470,000/ha) in coastal forests and from 22,000 to 111,000 per
acre (54,000-274,000/ha) in the interior [50].  Pure stands of western
redcedar may yield 60,704,168 seeds per acre (150,000,000/ha).  Poor
cone crops are rare [19].  Large seed crops occur every 3 to 4 years
[19,51].

Western redcedar seeds are small, 203,000 to 592,000 seeds per pound
(448,000-1,305/Kg) [8,50].  The seeds are dispersed primarily by wind.
However, the seeds have small wings and are not carried more than 400
feet (122 m) from the parent tree [8,48,50].

Germination:  Germination is epigeal.  Western redcedar seeds germinate
well without stratification and remain viable for at least 7 years
stored dry (5 to 8 percent moisture) at 0 degrees Fahrenheit (-18 deg C)
[50].  Stratification may improve the germination of some dormant seed
lots.  However, in others it may lower the germination capacity [51].
Haig [34] reported germination rates of 73 percent, and Schopmeyer [69]
reported germination rates of 34 to 90 percent.

Mineral soil has been found to be a better seedbed in many environments
than moss or duff, which may dry out rapidly [19,21,50].  Heavily shaded
seedbeds have been associated with the best germination of western
redcedar in British Columbia [19].  Rotten wood that is in contact with
the soil is the preferred seedbed in old western redcedar groves [50].
Graham [26] found that germination was best on burned surfaces.
 
Seedling development:  Western redcedar seedling survival is low [8,50].
Drought and high soil temperatures damage seedlings grown in full
sunlight [8,40].  Fungi, birds, insects, and smothering by fallen leaves
of deciduous shrubs are some other causes for the high mortality of
western redcedar seedlings [8,19].  Seedlings grow best in partial
shade, although they may fail on heavily shaded sites due to poor root
penetration [19].  Seedlings show high resistance to root flooding [47]
and respond well to removal of competition [57].  In one study, removal
of shrubs resulted in an increase in height growth of western redcedar
compared to unreleased trees [26].

Of all conifers in the northern Rocky Mountains, western redcedar and
western hemlock seedlings grow the slowest.  Annual height growth of
western redcedar seedlings is highly variable, from less than 0.39
inches (1 cm) in dense stands to over 7.5 inches (19 cm) in thinned
stands [26].

Vegetative reproduction:  Communities with closed canopies favor
vegetative reproduction over sexual reproduction [32].  Western redcedar
generally relies on vegetative reproduction in climax old-growth stands
with high soil moisture throughout the growing season [21,27].  The
frequent absence of adequate moisture in the upper soil layers of
well-drained sites often is responsible for western redcedar's reduced
ability to vegetatively reproduce on upland sites [28].  Three natural
types of vegetative reproduction occur:  (1) layering, (2) rooting of
fallen, living branches that have been torn off by wind or snow and have
fallen on wet soil; and (3) rooting along the trunks of fallen, living
trees [19,32,58].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Regional Distribution in the Western United States

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This species can be found in the following regions of the western United States (according to the Bureau of Land Management classification of Physiographic Regions of the western United States):

    1  Northern Pacific Border
    2  Cascade Mountains
    5  Columbia Plateau
    8  Northern Rocky Mountains
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Successional Status

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More info for the terms: climax, competition, forest, succession, tree

Obligate Climax Species

Western redcedar is very shade tolerant [8,21,50,51].  It is one of the
most shade tolerant species growing in cedar-hemlock ecosystems of the
northern Rocky Mountains [25].  It is usually considered a climax or
near climax species, but it can be found in all stages of forest
succession.  It invades disturbed areas as widely distributed seeds but
regenerates vegetatively in undisturbed areas, tolerating competition in
both [50].  Moisture and soil conditions strongly influence the
successional status of western redcedar.  It is climax on wet sites in
the Lake McDonald region of Glacier National Park and on calcium-rich
seepage habitats in British Columbia [51].  In Glacier National Park,
western redcedar enters pioneer communities.  The seedlings develop
rapidly in open stands of lodgepole pine (Pinus contorta) and western
larch (Larix occidentalis)[32].  It can survive as a late-seral or
coclimax tree on western-hemlock-dominated sites [21].  In Idaho,
western white pine (P. monticola) stands are slowly replaced by a
western hemlock-western redcedar climax [51].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Taxonomy

provided by Fire Effects Information System Plants
The currently accepted scientific name for western redcedar is Thuja
plicata Donn ex D. Don [8,50,51,57]. It is a member of the Cypress
family (Cupressaceae). Western redcedar hybridizes with Thuja
standishii. Hybrids are resistant to the leaf blight caused by
Didymascella thujina [50]. There are no recognized subspecies,
varieties, or forms.
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Value for rehabilitation of disturbed sites

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More info for the term: natural

Western redcedar can be planted on disturbed sites within its natural
range.  The erosion-control potential and long-term revegetation
potential of western redcedar have been rated as medium [13].  Western
redcedar may be the species of choice for reforesting high, brush-risk
areas near the coast [50].  It is suitable for planting on slightly dry
to wet nutrient-poor to nutrient-rich sites [commonly with Douglas-fir,
Sitka spruce (Picea sitchensis), Alaska-cedar (Chamaecyparis
nootkatensis), or western hemlock].  Western redcedar does best when
planted in mineral soils on upland sites and in well-decomposed organic
material on lowland sites [38].  Containerized western redcedar appears
to perform somewhat better than bareroot stock [26].  Direct seeding is
practical and effective where a mineral soil seedbed is available.
Methods for collecting, storing, and planting western redcedar seeds and
seedlings have been detailed [50,51,69].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Wood Products Value

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Western redcedar is an important commercial species throughout much of
its natural range [80].  In the Rocky Mountains, western redcedar
occupies some of the most productive sites, often producing stands with
high volume [26].  The wood is low in strength and soft but is very
resistant to decay, making it best suited for use as exposed building
material such as shingles, shakes, and exterior siding [57,69].
Hand-split western redcedar shakes sell for several times the price of
asphalt shingles but will last 100 years on a roof [8].  The wood is
fine and straight grained, which makes it suitable for interior
finishing [57].  Western redcedar wood is also used for utility poles,
fence posts, light construction pulp, clothes closets and chests, boats,
canoes, fish trap floats, caskets, crates, and boxes [50,80].
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Tesky, Julie L. 1992. Thuja plicata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Associated Forest Cover

provided by Silvics of North America
Pure stands of western redcedar cover some small areas, but it is usually associated with other tree species. Along the coast these include black cottonwood (Populus trichocarpa), bigleaf maple (Acer macrophyllum), western hemlock, mountain hemlock (Tsuga mertensiana), Sitka spruce, western white pine (Pinus monticola), lodgepole (shore) pine (P. contorta), Port-Orford-cedar (Chamaecyparis lawsoniana), Alaska-cedar (C. nootkatensis), incense-cedar (Libocedrus decurrens), Douglas-fir, grand fir, Pacific silver fir (Abies amabilis), red alder (Alnus rubra), Pacific madrone (Arbutus menziesii), and Pacific yew (Taxus brevifolia). Several of these species (black cottonwood, western hemlock, western white pine, Douglas-fir, grand fir, and Pacific yew) are also associated with western redcedar in the interior. Subalpine fir (Abies lasiocarpa), western larch (Larix occidentalis), Engelmann spruce (Picea engelmannii), white spruce (P. glauca), lodgepole pine, and ponderosa pine are also associated with redcedar in the interior.

Redcedar is a major component of two forest cover types (11): Western Redcedar (Society of American Foresters Type 228) and Western Redcedar-Western Hemlock (Type 227). It is a minor component of the following types:

210  Interior Douglas-Fir
212  Western Larch
213  Grand Fir
215  Western White Pine
218  Lodgepole Pine
221  Red Alder
222  Black Cottonwood-Willow
223  Sitka Spruce
224  Western Hemlock
225  Western Hemlock-Sitka Spruce
226  Coastal True Fir
229  Pacific Douglas-Fir
230  Douglas-Fir-Western Hemlock
231  Port-Orford-Cedar
232  Redwood

Some associated shrub species are listed in table 1. Several occur in both interior and coastal environments, but Rocky Mountain honeysuckle (Lonicera utahensis) and clematis (Clematis columbiana) are associated with redcedar only in the interior, whereas salmonberry (Rubus spectabilis) and red huckleberry (Vaccinium parvifolium) are found only on the Pacific slope. Pacific rhododendron (Rhododendron macrophyllum) is an abundant associate in coastal California, Oregon, and Washington, but it is rare and confined to isolated locations in south-coastal British Columbia. Salal (Gaultheria shallon) also is an abundant associate. Its range extends farther south than that of redcedar, but the northern limits of salal are nearly the same as the northern limits of western redcedar in coastal Alaska.

Table 1- Shrub species often associated with western redcedar in coastal and interior portions of its native range. Coastal Interior Both Coastal and Interior Oregongrape Mountian alder Western serviceberry   (Berberis nervosa)   (Alnus tenuifolia)   (Amelanchier alnifolia) Stink currant Creeping western barberry Rustyleaf menziesia   (Ribes bracteosum)   (Berberis repens)   (Menziesia ferruginea) Alaska blueberry Common juniper Thimbleberry   (Vaccinium alaskaense)   (Juniperus communis)   (Rubus parviflora) Box blueberry Cascade azalea Oceanspray   (Vaccinium ovatum)   (Rhododendron albiflorum)   (Holodiscus discolor) Red huckleberry Red raspberry Devilsclub   (Vaccinium parvifolium)   (Rubus idaeus)   (Oploponax horridum) Salmonberry Blue huckleberry Pachistima   (Rubus spectabilis)   (Vaccinium globulare)   (Pachistima myrsinites) Pacific rhododendron Rocky Mountain honeysuckle Baldhip rose   (Rhododendron macrophyllum)   (Lonicera utahensis)   (Rosa gymnocarpa) Salal Clematis Common snowberry   (Gaultheria shallon)   (Clematis columbiana)   (Symphoricarpos albus) Ovalleaf huckleberry   (Vaccinium ovalifolium) Some associated herb species are listed in table 2. Many are common in both coastal and interior environments. However, slough sedge (Carex obnupta) and Pacific water-parsley (Oenanthe sarmentosa) are limited to moist habitats west of the Cascades, whereas wild sarsaparilla (Aralia nudicaulis) and goldthread (Coptis occidentalis) occur with redcedar only in the interior.

Table 2- Herb species often associated with western redcedar in coastal and interior portions of its native range. Coastal Interior Both Coastal and Interior Threeleaf anemone Goldthread Maidenhair fern   (Anemone deltoidea)   (Coptis occidentalis)   (Adiantum pedatum) Deerfern Everlasting corydalis Ladyfern   (Blechnum spicant)   (Corydalis sempervirens)   (Athyrium filix-femina) Slough sedge Malefern Western swordfern   (Carex obnupta)   (Dryopteris filix-mas)   (Polystichum munitum) Pacific water-parsley Largeleaf avens Mountain woodfern   (Oenanthe sarmentosa)   (Geum macrophyllum)   (Dryopteris austriaca) Oregon oxalis Roundleaf alumroot Prince's-pine   (Oxalis oregana)   (Heuchera cylindrica)   (Chimaphila umbellata) Dwarf blackberry Panicle bluebells Bunchberry   (Rubus lasiococcus)   (Mertensia paniculata)   (Cornus canadensis) Youth-on-age Pine drops Skunkcabbage   (Tolmiea menziesii)   (Pterospora andromedea)   (Lysichitum americanum) White inside-out-flower Green pyrola False Solomon's-seal   (Vancouveria hexandra)   (Pyrola chlorantha)   (Smilacina stellata) Evergreen violet Pacific trillium   (Viola sempervirens)   (Trillium ovatum) Coastal redcedar plant communities in British Columbia have been classified into 24 associations under 9 alliances in the Thuja-Rubus spectabilis order (35). Communities listed in less detailed coastal classifications include redcedar/swordfern, redcedar/devilsclub/maidenhair fern, redcedar/maidenhair fern-ladyfern, redcedar-western hemlock/devilsclub/ladyfern, redcedar-grand fir/mountain boxwood, redcedar-grand fir/swordfern, redcedar-Douglas-fir/Oregongrape, redcedar-Sitka spruce-red alder/skunkcabbage-slough sedge, and redcedar/skunkcabbage (12,28).

The redcedar/skunkcabbage plant community also occurs in the interior; where redcedar/devilsclub, redcedar/devilsclub/coolwort (Tiarella trifoliata), redcedar/queenscup (Clintonia uniflora), and redcedar/maidenhair fern are found (7,28,42).

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Climate

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Coastal western redcedar receives from less than 890 mm (35 in) of annual precipitation to more than 6600 mm (260 in), mostly as winter rainfall; interior western redcedar, about 710 mm (28 in) in the north, 810 to 1240 mm (32 to 49 in) farther south-about half as spring and autumn rain, one-third as winter snow (2).

Although western redcedar is abundant in many forested swamps, it is sometimes found on sites that are too dry for western hemlock (Tsuga heterophylla) (12,39,42), probably because the root penetration of the redcedar is better. Redcedar leaves are not protected from excessive transpiration by cutin and wax.

Trees tolerate stagnant winter water tables averaging less than 15 cm (6 in) below the soil surface on the Olympic Peninsula (32). The species dominates wet ravines and poorly drained depressions in both Glacier National Park in Montana and the Selway Bitterroot Wilderness in Idaho (20,21).

Where sufficient precipitation is present, low temperature appears to limit the species' range. Length of the frost-free period abruptly decreases just above the tree's upper elevation limits on Vancouver Island. The northern limits of western redcedar lie between the 11.1° and 11.7° C (52° and 53° F) mean summer temperature isotherms in southeastern Alaska. Absolute minimum temperatures experienced by western redcedar in British Columbia are -10° to -30° C (14° to -22° F) in coastal populations, -14° to -47° C (7° to -53° F) in the interior (28). Western redcedar has a growing season of at least 120 frost-free days along the coast but as few as 75 frost-free days in some portions of its interior range. It is not resistant to frost and is sometimes damaged by freezing temperatures in late spring or early autumn.

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Damaging Agents

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Western redcedar is less susceptible than its associates to most damaging agents, but, as it is longer lived, damaged trees are common. Although they are as windfirm as Douglas-fir on dry sites, redcedars, are often windthrown in wet environments and are not resistant to windthrow on the moist sites where growth and yield are highest. Fire resistance also varies with environment. Redcedar is more severely damaged by fire than any of its associates along the coast but is less susceptible than Engelmann spruce, western hemlock, and subalpine fir in the interior (30).

Western redcedar suffers little damage from insects, but it is a host for several economically important insect species (13). One of the most important is the gall midge, Mayetiola thujae, which sometimes seriously damages redcedar seeds in Oregon, Washington, and British Columbia. Newly planted seedlings are occasionally damaged by a weevil (Steremnius carinatus) in British Columbia, and larger trees are killed by a bark beetle (Phloeosinus sequoiae) on poor sites in southeastern Alaska. The western cedar borer (Trachykele blondeli) causes degrade and cull in sawtimber.

More than 200 fungi are found on western redcedar, but it is less susceptible to pathological attacks than are most of its associates. Indeed, redcedar may be a suitable alternative species on coastal Douglas-fir and western hemlock sites where soils are infected with Phellinus weiri, Fomes annosus, or Armillaria spp.(37). Nevertheless, many attacks occur during the long lives of some redcedar trees, and the heartwood extractives that provide decay resistance are eventually detoxified through biodegradation by a series of invading fungi (25). As a result, the volume of accumulated decay in living trees is greater for western redcedar than for any other major conifer in British Columbia (25), and hollow old trees are common in the interior (7).

The major seedling disease, Didymascella thujina, is a leaf blight that infects 2nd- and 3rd-year nursery seedlings. As much as 97 percent of the natural redcedar reproduction may also be killed when this blight reaches epidemic proportions (2). Epidemics are rare in North America, however, and Didymascella is not as damaging here as it is in Europe, where the disease seriously limited production of planting stock until cycloheximide fungicides were developed (3,52).

In North America, the most important fungi attacking redcedar are root, butt, and trunk rots (23). The root and butt rots include Phellinus weiri, Armillaria mellea, and Poria subacida. Poria asiatica and P. albipellucida are the most important trunk rots near the coast; P. asiatica and Phellinus weiri are most important in the interior range of western redcedar (2). These rots are most evident in old stands, where much of the standing volume is often defective and unmerchantable.

Redcedar seedlings and saplings are often severely browsed by deer, elk, or rodents, and browse damage may be the most important stand-establishment problem (6).

Western redcedar is damaged more than Sitka spruce by salt spray. Its foliage is more severely damaged by sulfur dioxide than is the foliage of Douglas-fir, western hemlock, and Sitka spruce and less damaged than the foliage of subalpine fir and grand fir. Redcedar is damaged less than Douglas-fir by airborne fluorides and ozone (30).

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Flowering and Fruiting

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When grown in the open, western redcedars begin to produce strobili at 10 years of age and usually every other year thereafter. Strobilus development can be artificially induced at younger ages and increased in mature trees by girdling or treating with gibberellin (8). The species is monoecious; male and female strobili are produced on different branches of the same tree, at different heights-the reddish male strobili on lower branches and the green female strobili nearer the treetops and farther from the trunk (38,52). Anthesis and pollination occur during March and April in southern stands near the coast. They occur during May and June in coastal Alaska and interior stands (47).

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Genetics

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Population Differences Western redcedar seems to vary less than other northwestern conifer species. Lack of isoenzyme variation in newly germinated seedlings from western Oregon and eastern and western Washington indicates that redcedar populations contain little genetic polymorphism (5). Isoenzyme variability is also low in British Columbia (56). Leaf oil terpene composition is similar in populations at both low and high elevations in British Columbia, Washington, Oregon, Idaho, and Montana; but small differences between coastal and interior populations were recently detected by discriminant analysis of the chemical data (53). Seedlings from different seed sources usually have remarkably similar forms and growth rates when grown in the same environment. Inland populations are more tolerant of frost than those from coastal populations, however (46), and plantation trials indicate that provenances from Alaska are inferior to those from Oregon and Idaho when grown in Poland (31). Seed orchards have been established in Denmark, where resistance to leaf blight and frost have been shown to be homozygously recessive (52).

Races Several horticultural varieties of western redcedar are grown in North America. They include atrovirens, fastigiata, and pendula. Haploid and triploid varieties have been studied in Germany (31).

Hybrids Thuja plicata x Thuja standishii hybrids are resistant to the leaf blight caused by Didymascella thujina (52).

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Growth and Yield

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Intermediate redcedars are distinguishable from dominants by age 5 on good sites. Codominants usually can be differentiated from dominants by age 25. Mean annual height growth of the dominants is 0.5 m (1.64 ft) in 40- to 60-year-old, pure, second-growth stands on moist sites in western Washington (36). Annual radial increments of 10 mm or even 20 mm (0.4 to 0.8 in) occur on the best moist sites in the south coast region of British Columbia (39).

Pure, even-aged stands can attain volumes comparable to pure Douglas-fir stands by age 50 on high-quality upland sites in western Washington (37), Plantations should be dense (about 2,470 trees per hectare or 1,000/acre), and intermediate crown classes should be removed in a light thinning to reduce side shade at about age 25 (22,36). Stands of 370 to 430 crop trees/ha (150 to 175 crop trees/acre) at time of harvest may allow maximum diameter growth without causing poor form (36). Maintaining a nearly closed canopy at all times will benefit form because open-grown redcedars tend to develop excessively large limbs and multiple tops. Faster growing trees of acceptable quality can be grown at wide spacings if their lower holes are pruned (50), but percentages of latewood decrease significantly (49).

Volumes of 379 to 825 m³/ha (5,418 to 11,782 ft³/acre) were measured in 40- to 60-year-old pure second-growth stands on moist sites in western Washington (36). A yield model on medium sites in British Columbia indicates yields of 70 m³/ha (1,000 ft³/acre) at age 40, 350 m³/ha (5,000 ft³/acre) at age 115, and 595 m³/ha (8,500 ft³/acre) at age 270; maximum current annual increment occurs at 82 years and maximum mean annual increment at 130 years (34).

In Great Britain, the cumulative volume produced by normal western redcedar stands on poor sites is 50 m³/ha (714 ft³/acre) at age 20 and 953 m³/ha (13,620 ft³/acre) at age 80. On good sites, cumulative volume produced is 232 m³/ha (3,315 ft³/acre) at age 20 and 1839 m³/ha (26,268 ft³/acre) at age 80. The average age of maximum mean annual increment is 72 on poor sites and 58 on good sites in these British stands (22). At ages 20 and 50, cumulative volume production is lower for western redcedar than for Douglas-fir and Sitka. spruce in Great Britain, but by age 80 the redcedar volume production is higher than that of Douglas-fir and spruce (45).

Growth is often much slower. Suppressed redcedar trees that are 200 years old but only 7.6 cm. (3 in) in d.b.h. and 7.6 m (25 ft) tall are not unusual. Survival for such long periods of suppression may be due to the ability of the species to produce new root growth in full shade. It may also be a result of frequent root grafting. Dominant trees often support growth of the root systems and lower boles of suppressed trees (9).

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Reaction to Competition

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Only Pacific silver fir, western hemlock, and Pacific yew are more tolerant of shade than western redcedar (30). Its relative tolerance may be higher in warm than in cool areas, but redcedar is very tolerant wherever it grows, and it may be "the species of choice" for reforesting high, brush-risk areas near the coast (6). Often present in all stages of forest succession, redcedar occupies pioneer, seral, and climax positions (39). Multiple attributes seem to be responsible-redcedar invades disturbed areas as widely distributed seeds but regenerates vegetatively in undisturbed areas, tolerating competition in both (26). Nevertheless, it is usually considered a climax or near climax species.

Western redcedar is best managed on moist sites characterized by the presence of ladyfern, queenscup, mountain woodfern, oakfern (Gymnocarpium dryopteris), or thimbleberry. On poorly drained sites of lower quality, fertilizing with nitrogen appears to benefit growth (37). Urea seems to be a better source of that nitrogen than ammonium nitrate (54). Redcedar can be grown in stands of mixed species where uneven-aged management is practiced or when redcedar poles are to be produced under normal even-aged management regimes. But pure stands are more suitable for the long rotations needed to produce large sawtimber, shingles, and shakes.

Where western redcedar is managed in mixed-species or uneven-aged stands, its excellent shade tolerance and long life should be considered. Redcedar is usually overtopped by Douglas-fir, grand fir, western hemlock, and western white pine. It tolerates understory conditions in mixed-species stands but often grows slowly there. In uneven-aged stands, western redcedar can maintain acceptable growth rates over long periods, but it should not be given excessive crown space. Thinning from above may release the redcedars in mixed-species stands; thinning from below is preferable in uneven-aged stands of western redcedar. In the northern Rocky Mountains, growth response to release is best on large, young redcedars with green-yellow foliage growing on northerly aspects (17). Redcedars probably should not be released when overtopped, however, because much of the increased growth after their release often occurs in large branches and a spreading crown rather than stem wood (37).

Most western redcedars are harvested by clearcutting the mixed-species stands in which they grow. Because of steep terrain, decay, and breakage, redcedar harvesting costs are high and lumber recovery is low (55). Redcedars should not be left as scattered seed trees, however; even those along clearcut margins may be lost to windthrow or exposure. Effects of slash-burning vary with site conditions, but low-impact spring burns tend to benefit the mycorrhizal colonization of seedlings (6).

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Rooting Habit

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Tree roots are extensive. Redcedars made up only 17 percent of the basal area but accounted for 82 percent of the root length in a mixed-species stand in northern Idaho (29). Tap roots are poorly defined or nonexistent, but fine roots develop a profuse, dense network.

Western redcedar roots usually are deeper than the roots of western hemlock but shallower than the roots of western larch, western white pine, grand fir, and Douglas-fir (30). The soils on which these species usually grow may be responsible, however, because western redcedar, western hemlock, and Douglas-fir trees of similar size growing on similar soils have roots that penetrate to similar depths and extend over similar areas (10). Shallow root systems are most frequent where soil bulk density is high. Redcedar roots cannot grow in dense soils penetrated by the roots of Douglas-fir, red alder, lodgepole pine, and Pacific silver fir (30). Redcedar root systems also tend to be shallower and less extensive on wet sites than they are on deep, moderately dry soils.

Where a thick duff layer is present, many redcedar roots lie in the duff rather than in the underlying soil. Root grafting is common (9). Western redcedar mycorrhizae are of the vesicular-arbuscular type, and redcedar seedlings are more responsive to mycorrhizal inoculation than are the seedlings of redwood (Sequoia sempervirens), incense-cedar (Libocedrus decurrens), and giant sequoia (Sequoiadendron giganteum) (27).

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Seed Production and Dissemination

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Each mature strobilus usually produces only 3 to 6 seeds (8), but the strobili are often numerous and heavy seed crops are common. In dry years, conebearing stands in the interior tend to be on high, moist sites (14). Average annual seed crops vary from 247,000 to 2,470,000 seeds per hectare (100,000 to 1 million/acre) in coastal forests and from 54,000 to 274,000/ha (22,000 to 111,000/acre) in the interior. Major seedfall occurs during October and November in both ranges (2). Gibberellin-treated, containerized seed orchards should permit efficient management of seed production and harvesting (8).

Seeds are small- 448,000 to 1,305,000/kg (203,000 to 592,000/1b) (47). They fall faster and do not fly as far as the seeds of western hemlock, Sitka spruce, and Douglas-fir, but dissemination is adequate within 100 m (330 ft) of a seed source (4,30). The seeds usually germinate well without stratification, and they retain their initial viability for at least 7 years when stored dry (5 to 8 percent moisture) at -18° C (0° F) (8). Germination is epigeal.

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Seedling Development

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Most seeds escape rodent and bird predation, but seedling mortality is high during the germination period (15). Where moisture and temperature conditions are favorable, germination can occur in the autumn, winter, or spring. Almost no germination occurs after the first year, however. Seedbed quality may be critical.

Throughout the range of western redcedar, disturbed mineral soil seedbeds seem to be a major requirement for regeneration from seed (41). Although unburned soil benefits redcedar more than soil that has been scorched, slash burning favors redcedar by creating more mineral soil surfaces in cutover areas. Rotten wood that is in contact with the soil is the preferred seedbed in old redcedar groves (41). Partial shade is beneficial because drought and high soil temperature damage seedlings in full sunlight, and poor root penetration causes damage from drought in full shade (48).

Direct seeding in the autumn is successful where soil moisture is available, but large quantities of seed may be required to obtain adequate stocking. In the nursery, spring sowing is best; half-shaded seedbeds are recommended (47). Pelleting the seeds makes them more compatible with automated nursery sowing machinery (8). Containerized nursery seedlings can be produced in 7 months. They survive as well or better than bare-root stock when planted in coastal Oregon, Washington, and British Columbia, but 2-year-old bare-root stock tends to be most cost effective in the coastal range (6). When bare-root stock is planted, recently lifted dormant seedlings with low shoot/root ratios should be used and cold storage avoided whenever possible. Containerized stock planted in the spring appears to perform better than bare-root stock in the interior (18). Natural regeneration is important in the northern Rocky Mountains, where it is most frequently successful on westerly and northerly aspects in western redcedar habitat types (18).

Western redcedar seedlings are less tolerant of high soil temperature and of frost than are the seedlings of Engelmann spruce, grand fir, and Douglas-fir. The exposed upper foliage of young redcedars often sunburns severely (31). Roots of seedlings grow more slowly than the roots of Douglas-fir and incense-cedar, but they outgrow the roots of seedling western hemlock and Sitka spruce. Shoots have a longer growth period than any associated conifer. Non-rigid leaders are produced, and neither lateral nor terminal shoots form dormant buds. Lateral shoot growth is vigorous, amounting to at least 80 percent of terminal shoot growth in young redcedars (31). Seedlings account for most of the western redcedar regeneration in clearcuts and other disturbed areas. On good coastal sites, they grow as tall or taller than Douglas-fir, western hemlock, and Sitka. spruce seedlings during the first 5 years (6,51). The redcedars are subsequently overtaken by Douglas-fir (by age 10) and western hemlock (by age 15).

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Soils and Topography

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Western redcedar can tolerate a wide range of soil proper-ties in most localities. Soils on which it is most commonly found are in the orders Inceptisols, Ultisols, and Histosols. It is found on all landforms, soil textures, and parent materials on Vancouver Island (39). In southwestern Oregon, it grows on sedimentary, metasedimentary, gabbroic, dioritic, granodioritic, and occasionally even serpentinite and peridotite parent materials (24). Coarse sandy soils are not well suited to establishment and growth of redcedar in northern Idaho and northeastern Washington, but rocky slopes with limited soil development support redcedars in southeastern Alaska. Poorly drained organic soils also support redcedar south of Petersburg, AK. It grows on loams, clays, sands, chalk downland, and Molina-Juncus peat in England, but is most competitive on fine-textured lowland soils there. It grows well on shallow soils over chalk and can tolerate both acid and alkaline soil conditions (45).

Western redcedar seems able to survive and grow on soils that are low in nutrients and is found on such soils over much of its natural range. Site index is positively correlated with foliar nitrogen, sulfur, copper, boron, and chlorophyll. However, productivity may be improved by fertilization (44). When grown in well watered soil fertilized with nitrogen, phosphorus, and potassium, redcedar seedlings outgrow the seedlings of Douglas-fir (Pseudotsuga menziesii), grand fir (Abies grandis), Sitka spruce (Picea sitchensis), western hemlock, and ponderosa pine (Pinus ponderosa). Available nitrogen, calcium, and water appear to be the most important factors affecting growth and establishment of redcedar. Established redcedars tend to raise soil cation exchange capacities, pH's, and amounts of exchangeable calcium (1) and thus benefit the soils in which they grow.

Western redcedar grows from sea level to 910 m (3,000 ft) in southeastern Alaska. In British Columbia, the elevational range is higher-from sea level to 1190 m (3,900 ft). Redcedar is found in the interior from 320 m (1,050 ft) to 2130 m (7,000 ft).

The greatest range in elevation occurs in Oregon, where the species occurs from sea level to 2290 m (7,500 ft) at the rim of Crater Lake.

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Special Uses

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Shingles and shakes constitute the most important special use of redcedar. Attractive appearance, durability, lightness, and superior insulation qualities probably are responsible for its popularity as a roofing material. Wood is also used in utility poles, fenceposts, piling, paper pulp, clothes closets and chests, caskets, crates, boxes, beehives, and fishtrap floats. Perfumes, insecticides, medicinal preparations, veterinary soaps, shoe polishes, and deodorants are made from cedar leaf oil. Redcedar extractives and residues are used in lead refining, boiler-water additives, and glue extenders (31). When properly trimmed, redcedars make excellent hedges. Their drooping branches, thin fibrous bark, and flat sprays of scalelike leaves make redcedars attractive ornamental trees. The leaves are a major winter food for big game in the northern Rocky Mountains, and deer browse redcedar all year along the coast.

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Vegetative Reproduction

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Three types of natural vegetative reproduction occur: layering, rooting of fallen branches, and branch development on fallen trees. The resulting "veglings" are more abundant than seedlings in mature Idaho stands (40). Saplings that have been knocked down in the western Cascades often regenerate when their branches root.

Redcedar clones are easily propagated by the rooting of stem cuttings. Although untreated cuttings will root, a 1-minute dip in a 3,000 ppm solution or a 4-hour soak in a 200 to 400 ppm solution of indolebutyric acid improves rooting speed, the number of cuttings rooted, and the total length of roots per cutting. Ramets for seed orchards can be produced by treating cuttings with indolebutyric acid, then rooting them in a 1-to-1 mixture of peat and perlite (8). Young fragmented stems can be induced to bud after being soaked in a cytokinin solution, and the resulting buds can be rooted on a culture medium that contains napthalineacetic acid and kinetin (33).

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Distribution

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Western redcedar grows along the Pacific coast from Humboldt County, CA (lat. 40° 10' N.), to the northern and western shores of Sumner Strait in southeastern Alaska (lat. 56° 30' N.). In California, it is common only in the lower Mad River drainage and the wet region south of Ferndale in Humboldt County; it is found elsewhere only in isolated stands in boggy habitats (19). North of the California-Oregon border, the coastal range broadens to include the western slopes of the Cascade Range north of Crater Lake and the eastern slopes north of about latitude 44° 30' N. (12). Optimal growth and development of western redcedar are achieved near the latitudinal center of its range- Washington's Olympic Peninsula.

North of the Olympic Peninsula and Vancouver Island, the coastal range narrows again and is restricted to the Coast Ranges and offshore islands. A few scattered stands are found between the Coast Ranges and the Selkirk Mountains near the southern border of British Columbia, but redcedar's coastal range is essentially isolated from its interior range.

The interior range extends south from the western slope of the Continental Divide at latitude 54° 30' N. in British Columbia through the Selkirk Mountains into western Montana and northern Idaho (2). The southern limit is in Ravalli County, MT (lat. 45° 50' N.). With the possible exception of a few trees east of the Continental Divide near the upper end of St. Mary Lake, Glacier County, the eastern limit of the range of redcedar is near Lake McDonald in Glacier National Park, MT.


- The native range of western redcedar.

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Brief Summary

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Cupressaceae -- Cypress family

Don Minore

Western redcedar (Thuja plicata), also called Pacific redcedar, giant-cedar, arborvitae, canoe-cedar, and shinglewood, is the only Thuja species native to western North America. Extant redcedar volumes are estimated to be 824 million m³ (29 billion ft³) in British Columbia (43) and 228 million m³ (8 billion ft³) in the United States (16). Most of this volume is in mature trees, which have tapered, often-fluted bases, drooping branches, thin fibrous bark, and small scalelike leaves arrayed in flat sprays. Many have forked tops. They often reach ages of 800 to 1,000 years. One particularly large specimen in Washington has a d.b.h. of 592 cm (233 in), a height of 54.3 m (178 ft), and a crown spread of 16.5 m (54 ft). The wood is valuable and extensively used in a wide variety of products.

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Physical Description

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Tree, Evergreen, Monoecious, Habit erect, Trees without or rarely having knees, Tree with bark rough or scaly, Tree with bark shaggy or peeling, Young shoots in flat sprays, Buds not resinous, Leaves scale-like, Leaves opposite, Non-needle-like leaf margins entire, Leaf apex acute, Leaves < 5 cm long, Leaves < 10 cm long, Leaves not blue-green, Leaves white-striped, Scale leaves without raised glands, Scale leaf glands not ruptured, Scale leaves overlapping, Twigs glabrous, Twigs not viscid, Twigs without peg-like projections or large fascicles after needles fall, Berry-like cones orange, Woody seed cones < 5 cm long, Bracts of seed cone included, Seeds red, Seeds brown, Seeds winged, Seeds equally winged.
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Thuja plicata

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Thuja plicata is a large evergreen coniferous tree in the family Cupressaceae, native to the Pacific Northwest of North America. Its common name is western redcedar in the U.S.[2] or western red cedar in the UK,[3] and it is also called pacific red cedar, giant arborvitae, western arborvitae, just cedar, giant cedar, or shinglewood.[4] It is not a true cedar of the genus Cedrus. T. plicata is the largest species in the genus Thuja, growing up to 70 metres (230 ft) tall and 7 metres (23 ft) in diameter. It mostly grows in areas that experience a mild climate with plentiful rainfall, although it is sometimes present in drier areas on sites where water is available year-round, such as wet valley bottoms and mountain streamsides. The species is shade-tolerant and able to establish in forest understories and is thus considered a climax species. It is a very long-lived tree, with some specimens reaching ages of well over 1,000 years.

Indigenous peoples of the Pacific Northwest use the wood of this species for many purposes, such as building canoes, totem poles, and tools. The bark is harvested by indigenous peoples and processed into a fiber, which they use to make items such as rope, baskets, clothing, and rain hats. Because of its wide range of uses, the species is of great cultural importance to these people. Western redcedar wood is aromatic and rot-resistant and is used for applications such as the construction of shingles and siding. It has been introduced to cool temperate areas in other parts of the world, such as Northern Europe and New Zealand.

Description

Thuja plicata is a large to very large tree, ranging up to 45 to 70 metres (150 to 230 ft) tall and 2.4 to 7 m (8 to 23 ft) in trunk diameter,[5][6][7] larger than any other species in its genus.[8] The trunk swells at the base and has shallow roots.[5] The bark is thin, gray-brown, and fissured into vertical bands.[5] Trees growing in the open may have a crown that reaches the ground, whereas trees densely spaced together will exhibit a crown only at the top, where light can reach the leaves.[9] As the tree ages, the top is damaged by wind and replaced by inferior branches.[5] The species is long-lived; some trees can live well over a thousand years, with the oldest verified aged 1,460.[6][7]

The foliage forms flat sprays with scale-like leaves in opposite pairs, with successive pairs at 90 degrees to each other. The foliage sprays are green above and green marked with whitish stomatal bands below; they emit a strong aroma reminiscent of pineapple when crushed. The individual leaves are 1 to 4 millimetres (132 to 532 in) long and 1 to 2 mm (132 to 332 in) broad on most foliage sprays but up to 12 mm (12 in) long on strong-growing lead shoots.[6][7] The foliage of individual branchlets turns orange-brown before falling off in autumn.[5] Thuja plicata branches and foliage exhibit considerable morphological plasticity in response to light conditions. Branches growing in full sunlight produce denser foliage with more overlap, while shaded branches grow more horizontally, with less self-overlap.[10]

The cones are slender, 10 to 18 mm (38 to 1116 in) long, and 4 to 5 mm (532 to 316 in) broad, with 8 to 12 (rarely 14) thin, overlapping scales. They are green to yellow-green, ripening brown in fall about six months after pollination, and open at maturity to shed the seeds. The seeds are 4 to 5 mm (532 to 316 in) long and 1 mm (132 in) broad, with a narrow papery wing down each side. The pollen cones are 3 to 4 mm (18 to 532 in) long, red or purple at first, and shed yellow pollen in spring.[6][7]

Chemistry

The heartwood of western redcedar contains numerous chemical substances, such as plicatic acid, thujaplicatin methyl ether, hinokitiol and other thujaplicins, β-thujaplicinol, thujic acid, methyl thujate, 1,4-cineole, and γ-eudesmol.[11] Plicatic acid is believed to be the main irritant and contact allergen responsible for provoking allergic reactions and asthma exaggeration. This leads to occupational asthma in woodworkers that are exposed to western redcedar wood dust.[12] Thujaplicins serve as natural fungicides[13][14] which prevent the wood from rotting. This effect lasts around a century even after the tree is felled. However, thujaplicins are only found in older trees. Saplings do not produce the chemical, causing them to often develop rot at an early stage, causing some trees to grow with a somewhat hollow trunk, as the tree moves to heal itself as it grows.[15] Due to their fungicidal and anti-browning properties, thujaplicins are used in agriculture for fungal diseases and to prevent post-harvest decay.[16][17] Thujaplicins, like other tropolones, are potent chelating agents and bind divalent metal ions.[18] Basic and animal studies have shown that thujaplicins may have other biological properties, including antibacterial, antiviral, and antioxidant activities,[19] however reliable evidence on their effectiveness is still lacking.

Etymology

The species name plicata derives from the Latin word plicāre and means 'folded in plaits' or 'braided,' a reference to the pattern of its small leaves.[9]

Most authorities, both in Canada[20][21] and the United States[22][2][23][24] transliterate the English name in two words as 'western redcedar', or occasionally hyphenated as 'western red-cedar',[7] to indicate that it is not a true cedar (Cedrus), but it also appears as 'western red cedar' in some popular works. In the American horticultural trade, it is also known as the giant arborvitae, by comparison with arborvitae for its close relative Thuja occidentalis. Other names include giant red cedar, Pacific red cedar, shinglewood, British Columbia cedar (being the province's official tree),[5] canoe cedar, and red cedar.[6][15] Arborvitae comes from the Latin for 'tree of life'; coincidentally, Native Americans of the West Coast also address the species as "long life maker".[15]

One endonymous name for the tree is the Halkomelem word xepá:y,[25] from the roots xíp, meaning 'scratch' or 'line', and á:y, 'bark';[26] the former root may be in reference to both the lined or "folded/braided" appearance of the bark and the tree's ubiquity in carving and other forms of woodwork. It is called x̱ápay̓ay in the Squamish language.[27]

Taxonomy, evolution, and paleobiogeography

Thuja plicata is one of two Thuja species native to North America, the other being T. occidentalis. Nuclear genome analysis indicates that it is likely more closely related to T. koraiensis, which is native to the Korean Peninsula. The two species are hypothesized to have diverged in the Miocene.[28]

A 2008 study found that western redcedar populations in Oregon and California exhibit higher genetic variability than populations in the Rocky Mountains and coastal British Columbia, indicating that the species most likely colonized much of its current range from a single southern refugium after the Last Glacial Maximum.[29] Some studies have suggested the existence of an inland refugium during the Last Glacial Maximum, likely in central Idaho.[30][31] However, more recent genomic approaches corroborate the existence of only a single glacial refugium near the south of the current distribution, with subsequent expansion northward and inland since the last glacial maxium.[32] Refugial populations would have undergone repeated population bottlenecks during the glacial maxima of the Pleistocene.[30][32] Pollen cores indicate that Thuja plicata only became common in southern coastal British Columbia starting at approximately 6,000 YBP. This coincides with a transition from the warmer, drier climate of the early Holocene to a cooler, wetter climate. The species arrived later in the Canadian Rockies, as recently as 4,000–5,000 YBP.[33] Pollen Assemblages at Seeley Lake, on the eastern slope of the Coast Mountains east of Haida Gwaii, indicate that western redcedar arrived there at approximately 2,200 YBP.[34]

Distribution and habitat

A dense forest of western redcedar growing on flat terrain, with a lush understory
Thuja plicata often grows in moist valley bottoms.

Thuja plicata is among the most widespread trees in the Pacific Northwest. It is associated with Douglas-fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) in most places where it grows. It grows best on moist sites in areas with a wet, maritime climate,[35] and only occurs where precipitation exceeds 71 centimetres (28 in) annually.[22] It grows from the Cascade Range and Coast Mountains westward to the Pacific Ocean, from central South East Alaska (near the village of Kake) to northern California (growing closer to the coast at the northern and southern extremes). A disjunct population occurs inland from central-southeast British Columbia through the Idaho Panhandle.[5] The easternmost extent of its distribution occurs in Glacier National Park in Montana, where it is a major component of forests surrounding Lake McDonald.[36] Some small stands also occur just east of the continental divide in the park.[37] It is usually found from sea level to elevations of 1,100 m (3,600 ft),[5] but grows at altitudes of up to 2,290 m (7,510 ft) at Crater Lake in Oregon[22] and 1,500 m (4,900 ft) in Idaho.[5]

Western redcedar tolerates a wide variety of soil types.[22] It is present on all landforms and soil classes on Vancouver Island, but is restricted to wet, low-lying areas and streamsides in the drier eastern portions of its range.[38] It often grows in riparian areas throughout its range.[39] Unlike its associates western hemlock and Douglas-fir, western redcedar can grow in environments with stagnant groundwater present less than 5 centimetres (2.0 in) below the soil surface in the winter. It does not grow as well in areas with flowing groundwater, unlike another common associate, Sitka spruce (Picea sitchensis).[40] Western redcedars growing in the Rocky Mountains and along the eastern slope of the Washington Cascades are subject to higher mortality rates when growing on glacial till and sedimentary rocks than on other substrates. Western white pine (Pinus monticola) and Douglas-fir growing in these areas exhibit the opposite pattern, exhibiting low mortality rates on these substrates but high mortality rates on nutrient-poor metasedimentary rocks, a substrate on which western redcedar does not exhibit elevated mortality rates.[41]

Western redcedar is less cold-tolerant than many conifer species that it shares its range with and is vulnerable to frost damage in late spring and early fall. Its northern range limit in Southeast Alaska and its upper elevational limits on Vancouver Island are controlled by temperature, as moisture is plentiful in both areas and is therefore unlikely to be a limiting factor. In Southeast Alaska, it is absent from areas with mean summer temperatures less than 11 °C (52 °F). It is often absent from low-lying frost pockets in northern Idaho, where subalpine fir (Abies lasiocarpa) commonly grows instead. It does not grow in areas of coastal British Columbia with minimum temperatures of less than −30 °C (−22 °F), although some interior populations experience colder temperatures.[22]

It has been introduced to other temperate zones, including further north in Alaska, western Europe, Australia (at least as far north as Sydney), New Zealand,[42][43] and higher elevations of Hawaii.[44] The species was described as invasive in Great Britain by a 2004 survey,[45] although it is not listed as such by the United Kingdom Department for Environment, Food and Rural Affairs under the Wildlife and Countryside Act 1981 as of September 2022.[46] Its presence has also been recorded in Poland,[45] where it has been identified as a potentially problematic exotic species in the Białowieża Forest.[47]

Ecology

Use by wildlife

Western redcedar provides cover for bears, raccoons, skunks, and other animals which nest inside trunk cavities. It is used as a nest tree by cavity-nesting bird species such as yellow-bellied sapsuckers, hairy woodpeckers, tree swallows, chestnut-backed chickadees, and Vaux's swifts.[48] Pileated woodpeckers on the Olympic Peninsula prefer very large western redcedars with heart rot when selecting roosting sites, but do not use them for nesting, instead relying on Pacific silver fir (Abies amabilis) for that purpose.[49] Old-growth valley bottom forests of western redcedar and western hemlock are the preferred habitat for white-tailed deer in the Priest River drainage of northern Idaho. The dense canopies of these forests prevent deep snow accumulation, which hinders movement and results in increased energy expenditure for the deer.[50]

The foliage, especially that of saplings, is an important food source year-round for browsing ungulates such as Roosevelt elk and black-tailed deer, especially during the winter months when little else is available.[48] Western red-cedar foliage is more palatable for browsing herbivores than the commonly associated Sitka spruce and western hemlock. Browsing by introduced Sitka black-tailed deer is a major source of mortality for seedlings and saplings in old-growth forests on Haida Gwaii, where the deer lack natural predators. The presence of deer there has been linked to a decline in western redcedar recruitment.[51] The seeds may be eaten by birds and rodents such as deer mice, but are apparently not the preferred food source for most species, possibly due to their small size or unpleasant odor.[52][53] A 1937 study conducted in western Cowlitz County, Washington demonstrated a strong preference among deer mice for Douglas-fir and western hemlock seeds over western redcedar seeds.[54] However, pine siskins have displayed a preference for western redcedar seeds over those of Douglas-fir and western hemlock.[55]

Forest succession

Western redcedar appears in all stages of forest succession, but as one of the most shade-tolerant species in the Rocky Mountains and Pacific Northwest it is considered to be a climax species along with western hemlock.[56] It will readily establish and grow in the shade of other, less shade-tolerant species such as red alder (Alnus rubra), black cottonwood (Populus trichocarpa), or Douglas-fir, and prevent seedlings of those species from establishing themselves in its shade. However, western hemlock and Pacific silver fir are more tolerant of shade.[5]

A 2010 study found that Thuja plicata growing in old-growth forests 300 to 700 metres (980 to 2,300 ft) above sea level in the North Shore Mountains of British Columbia have higher growth rates than associated western hemlock and Pacific silver fir when growing underneath a closed canopy. The study also found that western redcedars do not increase their growth rates in response to canopy gap formation that occurs after the death of a mature overstory tree as much as the two other species. This result indicates that western redcedar may be less reliant on canopy gaps for recruitment than western hemlock and Pacific silver fir.[57] Western redcedar can also reproduce vegetatively via layering, as well as branch development from fallen branches or entire trees.[5][22]

On northeastern Vancouver Island, western redcedar grows along with western hemlock in relatively open-canopied forests with an understory dominated by salal (Gaultheria shallon). Another type of forest characterized by dense stands of western hemlock and Pacific silver fir with sparse understory vegetation also occurs in the area. The two forest types occur in areas with very similar environmental conditions and are separated by sharp boundaries, often less than 10 metres (33 ft) wide. Western redcedar recruitment is nearly absent in the western hemlock-Pacific silver fir forest type, and there is no evidence of a transitional stage between the two types. It has been hypothesized that, once established, these forest types are self-sustaining and are unlikely to change unless a major disturbance occurs.[58]

Fire ecology

It is considered to have low to moderate fire resistance, as its thin bark, shallow roots, low dense branching habit and flammable foliage confer little protection. Smaller trees are commonly killed by fire, but larger specimens often survive due to their size if they are not completely girdled. The intervals between fires within western redcedar stands tend to be very long, from 50 up to 350 years or more.[59]

Pathology

Western redcedar shows susceptibility of varying degrees to the following soil pathogens: Armillaria ostoyae, Fomitopsis pinicola, Heterobasidion annosum, Phaeolus schweinitzii, Phellinus weirii, Rhizinia undulata, and Postia sericeomollis.[60]

P. sericeomollis is responsible for brown cubical butt and pocket rot of cedar. It is the second-most common cause of decay in Western redcedar following P. weirii. Rather than forming a single column of decay in the heartwood, though, P. sericeomollis tends to cause rings or pockets of decay in the lower bole.[61]

While Western redcedar is a host to P. weirii, the fungus which causes the disease laminated root rot, it is rated as resistant while other conifers are rated as highly susceptible or susceptible.[62] P. weirii in Western redcedar expresses as a butt rot.[62]

In addition to P. weirii, western redcedar is also less susceptible to H. annosum and A. ostoyae than other conifer species.[63] Studies have found that western redcedar produces a phytochemical called thujaplicin which has been credited with granting the species its natural resistance to fungal attacks.[14] Additional research has shown that western redcedar responds to A. ostoye infection by producing necrophylactic periderms and resin ducts near infection sites, containing the spread of the fungus. Examination of co-occurring Douglas-fir and western hemlock demonstrated that western redcedar forms these structures in response to infection more frequently and successfully than the other two species.[64] Because of these natural defenses, it has been suggested that western redcedar may serve as a suitable alternative to other conifers when regenerating a site affected by these pathogens.[22]

Cedar leaf blight

Western redcedar foliage can become infected by the ascomycete fungus Didymascella thujina, which causes cedar leaf blight. The fungus causes leaves to turn brown and does not spread between adjacent leaves through hyphae. It is present throughout the native range of western redcedar, but reproduces best in relatively warm temperatures when western redcedar foliage is wet for long periods of time, with minimal wind exposure. Consequently, it is most common in wet coastal areas.[65] Significant variation in resistance to cedar leaf blight exists between populations of Thuja plicata; trees sourced from coastal populations exhibit higher resistance than trees from interior populations. Additional variation also occurs within populations.[66] A study published in 2013 projected a decrease in the incidence of cedar leaf blight among western redcedars in coastal British Columbia by late 21st century as a result of warmer, drier summers due to climate change. The authors of the study conceded that they were unable to incorporate the impacts of warmer winter temperatures into their model because research on the topic was lacking at the time of their study. They hypothesized that warmer winter temperatures could result in increased incidence of cedar leaf blight and cautioned that management decisions should take this uncertainty into account.[67]

Insect pathogens

Thuja plicata is a host to several destructive insect species such as the western cedar borer, cedar bark beetle, redcedar cone midge, and conifer seedling weevils.[68][48] Redcedar cone midge (Mayetiola thujae) larvae hatch from eggs laid between the scales of immature cones, then feed on the cone scales and seeds.[69] The application of insecticides has been used as a control measure.[70]

Cultivation

Like its relative Thuja occidentalis and many other conifer species, T. plicata is grown as an ornamental tree, and for screens and hedges, throughout the world in gardens and parks. A wide variety of forms, sizes, and colors is available.[71]

Cultivars

The following cultivars have gained the Royal Horticultural Society's Award of Garden Merit:

Uses

In indigenous societies

Klallam people and canoe, ca. 1914

Western redcedar is considered the Tree of Life by many of the First Nations of the Pacific Northwest, as the tree gave them everything that they needed for life - food, water (in the form of water tight woven cedar bark baskets), clothing, medicine, transportation (they made their canoes from the cedar tree), shelter (boards of wood from the cedar tree were used to build their long houses), and spirituality (in that the boughs were used in many ceremonies).[77] Some northwest coast tribes refer to themselves as "people of the Red Cedar" because of their extensive dependence on the tree for basic materials. The wood has been used for constructing housing and totem poles, and crafted into many objects, including masks, utensils, boxes, boards, instruments, canoes, vessels, houses, and ceremonial objects. Western redcedar is also associated with a long tradition of curing and cooking fish over the open fire. Roots and bark are used for baskets, bowls, ropes, clothing, blankets, and rings.[78][79]

A huge number of archaeological finds point to the continuous use of redcedar wood in native societies. Woodworking tools dating between 8,000 and 5,000 years ago, such as carved antlers, were discovered in shell middens at the Glenrose site, near Vancouver, British Columbia.[80] In Yuquot, on the west coast of Vancouver Island, tools dating 4,000 to 3,000 years old have been found.[80] The Musqueam site, also near Vancouver, yielded bark baskets woven in five different styles, along with ropes and ships dated to 3,000 years ago. At Pitt River, adzes and baskets were dated around 2,900 years ago. Wooden artefacts 1000 years old were unearthed on the east coast of Vancouver Island.[81]

Western redcedar was used extensively wherever it was found along the northwest coast (British Columbia, Washington state, parts of Alaska). Evidence of this use is found in CMTs (Culturally Modified Trees) that are found throughout the coast. When First Nations people removed the bark from cedars, it left a scar – which is considered a CMT. Other types of harvest (for planks, tinder, and other uses) leave different types of evidence of cultural modification.

A legend amongst the Coast Salish peoples describes the origins of the western redcedar. In this legend, there was a generous man who gave the people whatever they needed. When the Great Spirit saw this, he declared that when the generous man died, a great Red Cedar tree will grow where he is buried, and that the cedar will be useful to all the people, providing its roots for baskets, bark for clothing, and wood for shelter.[80]

Tools

The wood was worked primarily with the adze, which was preferred over all other tools, even ones introduced by Europeans. Alexander Walker, an ensign on the fur trade ship Captain Cook, reported that the indigenous peoples used an elbow adze, which they valued over tools brought by the Europeans, such as the saw or the axe, going so far as to modify traded tools back into an adze. Tools were generally made from stone, bone, obsidian, or a harder wood such as hemlock. A variety of hand mauls, wedges, chisels, and knives are also used.

Excavations done at Ozette, Washington, turned up iron tools nearly 800 years old, far before European contact. When James Cook passed the area, he observed that almost all tools were made of iron.[82] There has been speculation on the origin of these iron tools. Some theories include shipwrecks from East Asia or possible contact with iron-using cultures from Siberia, as hinted in the more advanced woodworking found in northern tribes such as the Tlingit.[82][83][84][85]

Wood

A totem pole outside a six-post house at the University of British Columbia

Harvesting western redcedar required some ceremony and included propitiation of the tree's spirits as well as those of the surrounding trees. In particular, many people specifically requested the tree and its brethren not to fall or drop heavy branches on the harvester,[86] a situation which is mentioned in a number of different stories of people who were not sufficiently careful. Some professional loggers of Native American descent have mentioned that they offer quiet or silent propitiations to trees which they fell, following in this tradition.

Felling of large trees such as western redcedar before the introduction of steel tools was a complex and time-consuming art. Typically the bark was removed around the base of the tree above the buttresses. Then some amount of cutting and splitting with stone adzes and mauls would be done, creating a wide triangular cut. The area above and below the cut would be covered with a mixture of wet moss and clay as a firebreak. Then the cut would be packed with tinder and small kindling and slowly burned. The process of cutting and burning would alternate until the tree was mostly penetrated through, and then careful tending of the fire would fell the tree in the best direction for handling. This process could take many days. A constant rotation of workers was involved to keep the fires burning through night and day, often in a remote and forbidding location.[87]

Once the tree was felled, the work had only just begun, as it then had to be stripped and dragged down to the shore. If the tree was to become canoes, then it would often be divided into sections and worked into rough canoe shapes before transport. If it were to be used for a totem pole or building materials, it would be towed in the round to the village.[88] Many trees are still felled in this traditional manner for use as totem poles and canoes, particularly by artists who feel that using modern tools is detrimental to the traditional spirit of the art. Non-traditionalists simply buy redcedar logs or lumber at mills or lumber yards, a practice that is commonly followed by most working in smaller sizes such as for masks and staves.

Because felling required such an extraordinary amount of work, if only planks for housing were needed, these would be split from the living tree. The bark was stripped and saved, and two cuts were made at the ends of the planking. Then wedges would be pounded in along the sides and the planks slowly split off the side of the tree.[89] Trees which have been so harvested are still visible in some places in the rainforest, with obvious chunks taken off of their sides. Such trees usually continue to grow perfectly well, since redcedar wood is resistant to decay. Planks are straightened by a variety of methods, including weighing them down with stones, lashing them together with rope, or forcing them between a line of stakes.[90]

Redcedar wood is used to make huge monoxyla canoes in which the men went out to high sea to harpoon whales and conduct trade.[91] One of those canoes, a 12-metre (38 ft) craft dug out about a century ago, was bought in 1901 by Captain John Voss, an adventurer. He gave her the name of Tilikum ('Relative' in Chinook jargon), rigged her, and led her in a hectic three-year voyage from British Columbia to London.[92]

Redcedar branches are very flexible and have good tensile strength. They were stripped and used as strong cords for fishing line, nets,[5] rope cores, twine, and other purposes where bark cord was not strong enough or might fray. Both the branches and bark rope have been replaced by modern fiber and nylon cordage among the aboriginal northwest coast peoples, though the bark is still in use for the other purposes mentioned above.

Bark

Illustration of women pulling bark from a tree, from Indian Legends of Vancouver Island by Alfred Carmichael

At the right time of year, the bark is easily removed from live trees in long strips. It is harvested for use in making mats, rope and cordage, basketry, rain hats, clothing, and other soft goods. The harvesting of bark must be done with care, as stripping too much bark will kill the tree. To prevent this, the harvester usually only harvests from trees that have not been stripped before.[93] After harvesting, the tree is not used for bark again, although it may later be felled for wood. Stripping bark is usually started with a series of cuts at the base of the tree above any buttresses, after which the bark is peeled upwards. To remove bark high up, a pair of platforms strung on rope around the tree are used and the harvester climbs by alternating between them for support. Since western redcedars lose their lower branches as all tall trees do in the rainforest, the harvester may climb 10 m (33 ft) or more into the tree by this method. The harvested bark is folded and carried in backpacks.[94] It can be stored for quite some time as mold does not grow on it and is moistened before unfolding and working. It is then split lengthwise into the required width and woven or twisted into shape. Bark harvesting was mostly done by women, despite the danger of climbing ten meters in the air, because they were the primary makers of bark goods.[95]

Today bark rope making is a lost art in many communities, although it is still practiced for decoration or art in a few places. Other uses of bark are still common for artistic or practical purposes. In recent years there has been a revival of cedar weaving in some communities, and along with it, new forms of cedar bark products. For example, in some recent weddings cedar roses are used to decorate the tables.

Timber

Canadian western redcedar cowl in the National Assembly for Wales

The soft red-brown timber has a tight, straight grain and few knots. It is valued for its distinct appearance, aroma, and its high natural resistance to decay, being extensively used for outdoor construction in the form of posts, decking, shingles, and siding.[96] It is commonly used for the framing and longwood in lightweight sail boats and kayaks. In larger boats it is often used in sandwich construction between two layers of epoxy resin and/or fiberglass or similar products. Due to its light weight – 390 to 400 kg/m3 (24 to 25 lb/cu ft) dried – it is about 30% lighter than common boat building woods, such as mahogany. For its weight it is quite strong but can be brittle. It glues well with epoxy resin or resorcinol adhesive.

Its light weight, strength, and dark, warm sound make it a popular choice for guitar soundboards, particularly among European guitar builders such as Lowden and Furch.

Western redcedar wood is export-restricted in the United States.[97] The tree is highly allergenic and woodworkers or loggers who work with it may have adverse reactions, including the development of occupational asthma, exacerbation of existing asthma, reduction of lung function, and eye irritation. Approximately 5% of workers are allergic to western redcedar. The US Occupational Safety and Health Administration has set a permissible exposure limit for western redcedar dust of 2.5 mg/m3 as a time-weighted average over eight hours.[98]

Essential oil

The essential oil of western redcedar leaves contains natural compounds, such as α-thujone, β-thujone, fenchone, sabinene, terpinen-4-ol and beyerene,[99] which have also been isolated from different other essential oils. Some of these substances are aroma compounds and are used in perfumery.[100] Thujones are GABAA receptor competitive antagonists, but do not have any pharmacological use due to their high toxicity and convulsive activity.[101]

Other uses

It is also widely used throughout Europe and America for making beehive components.

Its bark has been studied for applications in polyurethane.[102]

Used in the construction of windows and doors (joinery grade timber).

Notable specimens

The Quinault Lake Redcedar was the world's largest western redcedar.

The largest living specimen is the Cheewhat Giant, in Pacific Rim National Park Reserve on Vancouver Island, at 450 cubic metres (15,870 cu ft).[103] The tallest known individual is the Willaby Creek Tree south of Lake Quinault, 59 m (195 ft) in height.[104] The 'Quinault Lake Redcedar' was the largest known western redcedar in the world, with a wood volume of 500 m3 (17,650 cu ft). Located near the northwest shore of Lake Quinault north of Aberdeen, Washington, about 34 kilometres (21 mi) from the Pacific Ocean, it was one-third the volume of the largest known tree, a giant sequoia named 'General Sherman'. The Quinault Lake Redcedar was 53 m (174 ft) tall with a diameter of 5.9 m (19.5 ft) at breast height. The Quinault Lake Redcedar was destroyed by a series of storms in 2014 and 2016 and is now only a stump.[6][105] The fifth-largest known was the Kalaloch Cedar in Olympic National Park, at 350 m3 (12,370 cu ft),[106] until it was destroyed by a storm in March 2014.[107]

A redcedar over 71 m (233 ft) tall, 4.5 m (15 ft) in diameter, and over 700 years old stood in Cathedral Grove on Vancouver Island before it was set on fire and destroyed by vandals in 1972. That tree now lies in "Giant's Grave", a self-dug 'grave' created by the force of its own impact.[108] A specimen measuring 5.5 m (18 ft) diameter and 54 m (177 ft) tall on the Giant Red Cedar National Recreation Trail in the Idaho Panhandle National Forests is designated the "Champion Tree of Idaho".[109]

The Giant Cedar Stump is an ancient redcedar turned roadside attraction in Snohomish County, Washington.[110]

See also

References

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Thuja plicata: Brief Summary

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Thuja plicata is a large evergreen coniferous tree in the family Cupressaceae, native to the Pacific Northwest of North America. Its common name is western redcedar in the U.S. or western red cedar in the UK, and it is also called pacific red cedar, giant arborvitae, western arborvitae, just cedar, giant cedar, or shinglewood. It is not a true cedar of the genus Cedrus. T. plicata is the largest species in the genus Thuja, growing up to 70 metres (230 ft) tall and 7 metres (23 ft) in diameter. It mostly grows in areas that experience a mild climate with plentiful rainfall, although it is sometimes present in drier areas on sites where water is available year-round, such as wet valley bottoms and mountain streamsides. The species is shade-tolerant and able to establish in forest understories and is thus considered a climax species. It is a very long-lived tree, with some specimens reaching ages of well over 1,000 years.

Indigenous peoples of the Pacific Northwest use the wood of this species for many purposes, such as building canoes, totem poles, and tools. The bark is harvested by indigenous peoples and processed into a fiber, which they use to make items such as rope, baskets, clothing, and rain hats. Because of its wide range of uses, the species is of great cultural importance to these people. Western redcedar wood is aromatic and rot-resistant and is used for applications such as the construction of shingles and siding. It has been introduced to cool temperate areas in other parts of the world, such as Northern Europe and New Zealand.

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